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CANNABACEAE HEMP FAMILY

Alan T. Whittemore, except as noted

Tree, shrub, or erect or twining per; dioecious or flowers staminate and bisexual, wind-pollinated; epidermis with stiff hairs, glandular or not; sap watery.
Leaf: petioled; simple, unlobed or palmately lobed or compound, all alternate or lower opposite.
Inflorescence: terminal or axillary, unisexual or with both male and bisexual flowers.
Flower: perianth parts 4–6, free or fused; stamens 0 or 4–6; ovary 0 or 1, superior, chamber 1, ovule 1, style 1, short, stigmas 2, slender, plumose.
Fruit: drupe or achene, occasionally ± enclosed in persistent perianth.
11 genera, ± 100 species: temperate, tropical areas worldwide. [Sytsma et al. 2002 Amer J Bot 89:1531–1546] —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Mitchell 1988 Bull New York State Mus Nat Hist 464:17–23]

Key to Cannabaceae

HUMULUS HOP

Alan T. Whittemore & Elizabeth McClintock

Annual, perennial herb, twining herbaceous vine, hairs occasionally forked.
Leaf: lower opposite, palmately 3–5-lobed, upper alternate, unlobed.
Staminate inflorescence: panicle, ± open.
Pistillate inflorescence: head- or cone-like, generally pendent ( erect).
Pistillate flower: perianth parts fused into short, unlobed tube or ring, persistent.
Fruit: enclosed in enlarged perianth subtended by papery bract.
2 species: n temperate. (Probably latinized from Low German: hop)

H. lupulus L. EUROPEAN HOP
NATURALIZED
Perennial.
Leaf: generally 3–5-lobed; blade ± cordate, coarsely serrate.
Pistillate inflorescence: 2.5–5 cm, ± oblong in fruit.
Uncommon. Disturbed places, persisting from cultivation; < 3000 m. Outer North Coast Ranges, Cascade Range, n&c Sierra Nevada, South Coast, Western Transverse Ranges, San Gabriel Mountains; native to Eurasia. [Humulus lupulus var. neomexicanus A. Nelson & Cockerell] Orn vine and cultivated as major flavor source for beer. Apr–Aug [Online Interchange]
Unabridged note: The following (and possibly other) accessions, if verified, would represent range extensions (as indicated): RSA462307 (s SNH).

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.