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CANNABACEAE HEMP FAMILY

Alan T. Whittemore, except as noted

Tree, shrub, or erect or twining perennial herb; dioecious or flowers staminate and bisexual, wind-pollinated; epidermis with stiff hairs, glandular or not; sap watery. Leaf: petioled; simple, unlobed or palmately lobed or compound, all alternate or lower opposite. Inflorescence: terminal or axillary, unisexual or with both male and bisexual flowers. Flower: perianth parts 4–6, free or fused; stamens 0 or 4–6; ovary 0 or 1, superior, chamber 1, ovule 1, style 1, short, stigmas 2, slender, plumose. Fruit: drupe or achene, occasionally ± enclosed in persistent perianth.
11 genera, ± 100 species: temperate, tropical areas worldwide. [Sytsma et al. 2002 Amer J Bot 89:1531–1546] —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Mitchell 1988 Bull New York State Mus Nat Hist 464:17–23]

Key to Cannabaceae

HUMULUS HOP

Alan T. Whittemore & Elizabeth McClintock

Annual, perennial herb, twining herbaceous vine, hairs occasionally forked. Leaf: lower opposite, palmately 3–5-lobed, upper alternate, unlobed. Staminate inflorescence: panicle, ± open. Pistillate inflorescence: head- or cone-like, generally pendent (erect). Pistillate flower: perianth parts fused into short, unlobed tube or ring, persistent. Fruit: enclosed in enlarged perianth subtended by papery bract.
2 species: northern temperate. (Probably latinized from Low German: hop)

H. lupulus L. EUROPEAN HOP
NATURALIZED
Perennial herb. Leaf: generally 3–5-lobed; blade ± cordate, coarsely serrate. Pistillate inflorescence: 2.5–5 cm, ± oblong in fruit.
Uncommon. Disturbed places, persisting from cultivation; < 3000 m. Outer North Coast Ranges, Cascade Range, n&c Sierra Nevada, South Coast, Western Transverse Ranges, San Gabriel Mountains; native to Eurasia. [Humulus lupulus var. neomexicanus A. Nelson & Cockerell] Ornamental vine and cultivated as major flavor source for beer. Apr–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Humulus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=28532, accessed on Aug 1 2014

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click for enlargement Humulus lupulus
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© 2002 Bjørnar Olsen

Bioregions in which Humulus lupulus occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.