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Key to families | Table of families and genera

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CAMPANULACEAE BELLFLOWER FAMILY

Nancy R. Morin, except as noted

Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, ± fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or ± fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1–3, placentas axile or parietal, ovules many, style generally 1, 2–5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
± 90 genera, ± 2500 species: worldwide. [Haberle et al. 2008 J Molec Evol 66:350–361] Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Lammers 2007 World Checklist and Bibliography of Campanulaceae. Royal Botanic Gardens, Kew.]

Key to Campanulaceae

HOWELLIA

1 sp. (Thomas (1842–1912) and Joseph (1830–1912) Howell, brothers and botanists in Pacific Northwest)

H. aquatilis A. Gray WATER HOWELLIA
NATIVE
Annual, aquatic, rooted, glabrous. Stem: submersed, ascending to erect, or floating; 10–60 cm. Leaf: alternate (sometimes ± opposite or whorled), cauline, linear, entire or minutely toothed, sessile. Inflorescence: flowers 1 in leaf or bract axils, scattered; pedicels stout, 1–4(8) mm; bract 1 per flower. Flower: inverted; sepals 1/6–2/5 ovary, narrowly triangular; corolla white, 0 in lower flowers or not, tube 2–2.7 mm, narrowly cylindric, split adaxially ± to base, ± 2-lipped, 2 adaxial lobes ± 1–1.3 mm, narrowly strap-shaped, spreading, 3 abaxial lobes ± 1–1.3 mm; stamens fused into tube, anthers 0.2–0.9 mm, 2 shorter anthers minutely appendaged; ovary inferior, 5–13 mm, ellipsoid, stigma head-like. Fruit: 5–13 mm, 1–2 mm diam, ellipsoid, 1-chambered, depressed-conic at top, open on sides by slits or irregular tears. Seed: 2–4 mm, cylindric, shiny, chestnut-brown.
Seasonal ponds; 1100–1500 m. High North Coast Ranges; to Washington, Montana. Jun–Aug [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 2 2014
Citation for this treatment: [Author of taxon treatment] 2013. Howellia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=28520, accessed on Sep 2 2014

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Geographic subdivisions indicated for the distribution of Howellia aquatilis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.