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Margriet Wetherwax, except as noted

Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, [1]2–4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.

Key to Plantaginaceae


1 sp.: southeastern California. (Arthur H. Holmgren (1912–1992), Noel H. Holmgren (b. 1937), Patricia K. Holmgren (b. 1940)) [Ghebrehiwet et al. 2000 Plant Syst Evol 220:223–239]
Unabridged references: [Elisens 1985 Syst Bot Monogr 5:1–97]

H. petrophila (Coville & C.V. Morton) Elisens ROCK LADY
Perennial herb, glandular-puberulent to -hairy. Stem: erect to pendent, densely branching, becoming woody at base. Leaf: generally alternate; petiole 12–27 mm, not twining; blade 12–35 mm, 14–27 mm wide, ± round to reniform, irregularly bristly-dentate; veins palmate. Inflorescence: flowers 1 in leaf axils; pedicel 1–4 mm. Flower: sepals 5, ± equal, 9–13 mm, irregularly bristly-dentate; corolla white to pale yellow, tube-throat 20–24 mm, sac-like extension at base absent, floor with 2 longitudinal yellow folds, lower lip base generally not swollen, generally not closing mouth; stamens included, lower filaments 7–9 mm, upper 12–14 mm; style 9–10 mm. Fruit: 8–10 mm, ± spheric; chamber 1, septum incomplete, dehiscent by 2–3 pores. Seed: many, 2–3 mm, pitted.
Limestone crevices of canyons; 700–1700 m. n Mojave Desert (Titus, Fall canyons, Death Valley region, Inyo Co.). [Maurandya petrophila Coville & C.V. Morton] Apr–Jun [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 25 2015
Citation for this treatment: [Author of taxon treatment] 2013. Holmgrenanthe, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 25 2015

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Geographic subdivisions indicated for the distribution of Holmgrenanthe petrophila Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.