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VERBENACEAE VERVAIN FAMILY

Dieter H. Wilken

Annual to shrub [tree], generally hairy. Leaf: cauline, opposite, generally toothed; stipules 0. Inflorescence: raceme, spike, or head, generally elongated in fruit; bract generally 1 per flower. Flower: bisexual; calyx generally 4–5-toothed; corolla 4–5-lobed, radial to bilateral, salverform to 2-lipped; stamens 4–5 (if 4, generally in unequal pairs), epipetalous; ovary superior, 2- or 4-lobed, generally 2- chambered, style 1, often with 2 unequal lobes, only 1 stigmatic, lateral. Fruit: 2 or 4 nutlets, drupe-like, or capsule.
± 31 genera, ± 920 species: especially America tropics. Some cultivated (Lantana, Verbena, Vitex); some weedy worldwide (Lantana); some used for wood (Tectona, teak). Avicennia included in Acanthaceae. [Marx et al. 2010 Amer J Bot 97:1647–1663] —Scientific Editor: Bruce G. Baldwin.

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 21 2014
Citation for this treatment: [Author of taxon treatment] 2013. Verbenaceae, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=281, accessed on Aug 21 2014

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