Jepson Herbarium
		University of California, Berkeley

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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |

Annual to shrub, or non-green root parasite, generally bristly or sharp-hairy.
Stem: prostrate to erect.
Leaf: cauline, often with basal rosette, simple or compound, generally alternate.
Inflorescence: cymes, generally elongate, panicle-, raceme-, or spike-like, generally coiled in flower (often described as scorpioid), generally uncoiled in fruit, or heads, spikes, or panicles, or flowers 1–2 per axil.
Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla generally (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages 0 or 5 at top of tube, alternate stamens; stamens epipetalous; ovary superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched.
Fruit: nutlets 1–4, free ( fused), smooth to roughened, prickly or bristly or not, or valvate or circumscissile capsule.
± 120 genera, ± 2300 species: tropics, temperate, especially w North America, Medit; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum). Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. [Olmsted et al. 2000 Molec Phylogen Evol 16:96–112] Recently treated to include Hydrophyllaceae, Lennoaceae. —Scientific Editors: Ronald B. Kelley, Robert Patterson, Bruce G. Baldwin, Thomas J. Rosatti.

Key to Boraginaceae

Perennial, scapose; root caudex-like.
Leaf: simple, in basal rosette, spreading or ascending; blade tapered to petiole, generally entire, generally ciliate.
Inflorescence: flowers 1; peduncle erect or spreading, 1–10 cm, slender.
Flower: calyx lobes 2–9 mm, generally unequal, glabrous to hairy, ciliate; corolla scales 0, tube generally dense-hairy inside, throat generally yellow, lobes glabrous to hairy, white or ± blue, generally tinged or marked with lavender or purple; stamens included, generally unequal, filament base widened; ovary hairy, chamber 1, style 1, stigmas 2, 2–5 mm.
Fruit: 5–11 mm, ovoid, hairy.
Seed: many, ovoid, angular, red-brown, honeycombed or pitted.
2 species: w US, n Mex. (Greek: evening or western centaur)

Rhizomes 0.
Leaf: generally > 6, < 8 cm, < 3 cm wide, oblanceolate to elliptic or ovate.
Inflorescence: flowers generally > 5.
Flower: corolla 10–30 mm, limb 10–20 mm wide, lobes 3–10 mm, oblong.
n=8. Wet meadows, flats, valleys; 770–2620 m. Klamath Ranges, High Cascade Range, High Sierra Nevada, Tehachapi Mountain Area, Western Transverse Ranges, San Bernardino Mountains, Great Basin Floristic Province, s Mojave Desert (Rabbit Springs, San Bernardino Co.);

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.