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APIACEAE (Umbelliferae) CARROT FAMILY

Lincoln Constance & Margriet Wetherwax, except as noted

Annual to perennial herb [ shrub, tree], generally from taproot.
Stem: generally ± scapose, generally ribbed, hollow.
Leaf: basal and generally cauline, generally alternate; stipules generally 0; petiole base generally sheathing stem; blade generally much dissected, occasionally compound.
Inflorescence: umbel or head, simple or compound, generally peduncled; bracts present in involucres or 0; bractlets generally present in " involucels".
Flower: many, small, generally bisexual (or some staminate), generally radial (or outer bilateral); calyx 0 or lobes 5, small; petals 5, free, generally ovate or spoon-shaped, generally incurved at tips, generally ± ephemeral; stamens 5; pistil 1, ovary inferior, 2-chambered, generally with a ± conic, persistent projection or platform at tip subtending 2 free styles.
Fruit: 2 dry, 1-seeded halves (= mericarps), separating from each other but generally ± persistent to central axis; ribs on halves 5, 2 marginal, 3 to back; oil tubes 1–several per interval between ribs.
300 genera, 3000 species: ± worldwide, especially temperate; many cultivated for food or spice (e.g., Carum, caraway; Daucus; Petroselinum); Bupleurum lancifolium Hornem. is historical garden weed; some toxic (e.g., Conium). Mature fruit generally critical in identification, shape given in outline. Hydrocotyle moved to Araliaceae. Petroselinum crispum (Mill.) A.W. Hill is a waif. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Apiaceae

HERACLEUM HOGWEED
Perennial from taproot or clustered roots.
Stem: stout, erect, generally branched, hollow.
Leaf: blades oblong to round, ternately, pinnately, or palmately compound ( simple), leaflets large, lobed or serrate; distal-most cauline generally reduced to large sheaths.
Inflorescence: umbels compound, large, margins generally sterile; bracts 0–few, generally deciduous; bractlets generally present, persistent; rays, pedicels many, spreading- ascending.
Flower: marginal bilateral, outer petals > others, 2-lobed; calyx lobes generally 0; petals wide, white, ± yellow.
Fruit: oblong-ovate to round or obcordate, compressed front-to-back; ribs unequal, marginal thin-winged, veined near outer margin, others thread-like; oil tubes 1–2 per rib-interval, unequal in length; fruit axis divided to base.
Seed: face flat.
± 80 species: Eurasia, e Africa, 1 in North America. (Hercules, presumably from large stature of some species)

H. maximum W. Bartram COW PARSNIP, GIANT HOGWEED
NATIVE
Plant 1–3 m, stout, tomentose, strong-scented.
Leaf: round to reniform; petioles 1–4 dm, widely sheathing, distal sheaths enlarged, bladeless; blades 2–5 dm wide, ternate, leaflets 1–4 dm wide, ovate to round, cordate, lobed, coarsely serrate.
Inflorescence: tomentose or long-hairy; peduncle 5–20 cm; rays 15–30, 5–10 cm, unequal; pedicels 8–20 mm.
Flower: petals obovate, white.
Fruit: 8–12 mm, obovate to obcordate, ± hairy.
2n=22. Moist places, wooded or open; < 2900 m. California Floristic Province, Great Basin Floristic Province; to Alaska, e United States, Arizona. [Heracleum lanatum Michx.] Only native Apiaceae sp. found from e to w North America. Apr–Jul [Online Interchange]
Unabridged note: Relationship to some Eurasian taxa unclear.

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Next taxon: Ligusticum

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.