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Indexes to all accepted names and synonyms:
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Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1–2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1–4, free (fused), smooth to roughened, prickly or bristly or not.
± 120 genera, ± 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. [Olmstead et al. 2000 Molec Phylogen Evol 16:96–112] Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized. —Scientific Editors: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Key to Boraginaceae
Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often ± woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally ± = blades, ± winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, ± bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin.Key to Hackelia
40 species: generally western North America, southeastern Asia. (J. Hackel, Czech botanist, 1783–1869) Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula.
Unabridged references: [Gentry & Carr 1976 Mem New York Bot Gard 26:121–227]
Unabridged note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators.
Stem: 4–8 dm; hairs generally spreading, ± dense below middle, ± dense to ± sparse above. Leaf: velvety-hairy; basal generally 5–17 cm, 0.5–2 cm wide, narrow-elliptic to oblanceolate; lower cauline reduced, ovate to narrow-lanceolate, ± clasping upward. Inflorescence: ± dense, branches several, few-flowered; pedicel 6–20 mm in fruit. Flower: calyx 2–3 mm; corolla tube generally >> calyx, generally ± purple, throat ± closed, limb 12–20 mm diam, generally blue to ± purple (pink), appendages long-exserted, much longer than wide, recurved, 2-pronged at tip, white; anthers hidden. Fruit: nutlets 4.5–6.5 mm, dull, roughened, abaxial prickles many, evenly distributed, ± = marginal.
Dry, open slopes, forest clearings, roadsides; 1350–2750 m. High Sierra Nevada; western Nevada. Jun–Aug [Online Interchange]
Unabridged synonyms: [Hackelia longituba I.M. Johnst.]
Unabridged note: Though corolla distinctive, mistaken for Hackelia nervosa, Hackelia mundula: the unique long-exserted recurved corolla appendages and overall velvety hairs distinguish it from Hackelia nervosa; the exserted corolla tube, white corolla appendages, and dull, roughened nutlets separate it from Hackelia mundula. A CalPhoto image taken in southern Sierra Nevada shows a pink flowered plant, no doubt indicating introgression with Hackelia mundula. The suspected occurrence of Hackelia velutina in High Cascade Range based on multiple CalPhoto images in the Lassen Volcanic National Park area, though it is uncertain whether voucher specimens exist.
Previous taxon: Hackelia sharsmithii
Next taxon: Harpagonella
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Hackelia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=27544, accessed on Dec 22 2014
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© 2009 Keir Morse
|Geographic subdivisions indicated for the distribution of Hackelia velutina|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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