Directory       News       Site Map       Home
    Jepson eFlora: Taxon page
Key to families | Table of families and genera

Previous taxon Indexes to all accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Previous taxon


Mihai Costea, family description, key to genera, revised by Thomas J. Rosatti & Elizabeth H. Zacharias

Annual to shrub; hairs simple, stellate, or glandular; plants in several genera scaly, mealy, or powdery from collapsed glands; monoecious, dioecious, with bisexual flowers, or with both bisexual and unisexual flowers. Stem: occasionally fleshy. Leaf: blade simple, generally alternate, occasionally fleshy or reduced to scales, veins pinnate; stipules 0. Inflorescence: raceme, spike, catkin-like, spheric head, axillary clusters of flowers, or flowers 1; bracts 0–5, herbaceous, generally persistent or strongly modified in fruit, wings, tubercles or spines present or 0. Flower: bisexual or unisexual, small, generally green; calyx parts (1)3–5, or 0 in pistillate flowers, free or fused basally (or ± throughout), leaf-like in texture, membranous, or fleshy, deciduous or not, often strongly modified in fruit; corolla 0; stamens 1–5, opposite sepals, filaments free, equal; anthers 4-chambered; ovary superior (1/2-inferior), chamber 1; ovule 1; styles, stigmas 1–4 (or stigmas sessile). Fruit: achene or utricle, generally falling with persistent calyx or bracts. Seed: 1, small, lenticular to spheric; seed coat smooth to finely dotted, warty, net-like, or prickly, margin occasionally winged.
100 genera, 1500 species: worldwide, especially deserts, saline or alkaline soils; some cultivated for food (Beta vulgaris subsp. vulgaris, beet, Swiss chard; Spinacia oleracea L., spinach; Chenopodium quinoa Willd., quinoa); and some worldwide, naturalized ruderal or noxious agricultural weeds. Nitrophila treated in Amaranthaceae, Sarcobatus treated in Sarcobataceae. Key to genera revised by Elizabeth H. Zacharias to incorporate Extriplex and Stutzia, 2 genera segregated from Atriplex. —Scientific Editors: Bruce G. Baldwin, David J. Keil, Thomas J. Rosatti.

Key to Chenopodiaceae


Margriet Wetherwax & Dieter H. Wilken

1 sp. (Asa Gray, eminent American botanist, Harvard University, 1810–1888) [Holmgren 2003 FNANM 4:306–307]

G. spinosa (Hook.) Moq.
Shrub rounded; scaly-puberulent, hairs branched, glabrous in age; generally dioecious. Stem: generally 3–10(15) dm, branches many, stiff; bark red-brown, ± white-ribbed, peeling in strips, older bark gray; twigs spine-like in age. Leaf: alternate, 5–25(40) mm, generally spoon-shaped to oblanceolate, flat, entire, tapered to short-petioled, blade green, tip generally ± white. Staminate inflorescence: spike-like, terminal, 7–18 mm; bract ± leaf-like; flowers 2–5 per cluster. Pistillate inflorescence: ± spike-like, axillary or terminal, 6–18 cm in fruit; flowers 1–few per cluster; bracts 3–10 mm, ± leaf-like; fruit bracts 2, 7–15 mm, fused, together sac-like, ± round, flat, winged, white to red-tinged, margins entire. Staminate flower: calyx lobes 4, 1.5–2 mm, enclosing stamens; stamens 4–5. Pistillate flower: stigmas 2, exserted. Fruit: generally 1.5–2 mm, brown.
2n=36. Sandy to gravelly soils in scrub, pinyon/juniper woodland; 300–2900 m. High Sierra Nevada (e slope), Tehachapi Mountain Area, se San Joaquin Valley, Western Transverse Ranges (n slope), Great Basin Floristic Province, Mojave Desert, nw Sonoran Desert; to Washington, Montana, New Mexico. Mar–Jun [Online Interchange]
Unabridged synonyms: [Chenopodium spinosum Hook.]

Previous taxon: Grayia
Next taxon: Halogeton


Name search

Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Mar 30 2015
Citation for this treatment: [Author of taxon treatment] 2013. Grayia, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Mar 30 2015

Copyright © 2014 Regents of the University of California
We encourage links to these pages, but the content may not be downloaded for reposting, repackaging, redistributing, or sale in any form, without written permission from The Jepson Herbarium.

Geographic subdivisions indicated for the distribution of Grayia spinosa Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
View all CCH records


CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.