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FABACEAE (Leguminosae) LEGUME FAMILY

Martin F. Wojciechowski, except as noted

Annual to tree.
Leaf: generally alternate, generally compound, generally stipuled, generally entire, pinnately veined
Inflorescence: generally raceme, spike, umbel or head; or flowers 1–few in axils.
Flower: generally bisexual, generally bilateral; hypanthium 0 or flat to tubular; sepals generally 5, generally fused; petals generally 5, free, fused, or lower 2 ± united into keel (see 3, Key to Groups, for banner, wings); stamens 10 or many (or [1], 5, 6, 7, 9), free or fused or 10 with 9 filaments at least partly fused, 1 (uppermost) free; pistil 1, ovary superior, generally 1-chambered, ovules 1–many, style, stigma 1.
Fruit: legume, including a stalk-like base (above receptacle) or not.
Seed: 1–many, often ± reniform, generally hard, smooth.
± 730 genera, 19400 species: worldwide; with grasses, requisite in agriculture, most natural ecosystems. Many cultivated, most importantly Arachis, peanut; Glycine, soybean; Phaseolus, beans; Medicago, alfalfa; Trifolium, clovers; many orns. [Lewis et al. (eds) 2005 Legumes of the World. RBG, Kew] Unless stated otherwise, fruit length incl stalk-like base, number of 2° leaflets is per 1° leaflet. Upper suture of fruit adaxial, lower abaxial. Anthyllis vulneraria L. evidently a waif, a contaminant of legume seed from Europe. Laburnum anagyroides Medik., collected on Mount St. Helena in 1987, may be naturalized. Ceratonia siliqua L., carob tree (Group 2), differs from Gleditsia triacanthos L. in having evergreen (vs deciduous) leaves that are 1-pinnate (vs 1-pinnate on spurs on old stems, 2-pinnate on new stems) with 2–5(8) (vs 7–17) 1° leaflets, commonly cult, now naturalized in s CA. Aeschynomene rudis Benth. <Noxious weed>, Halimodendron halodendron (Pall.) Voss <Noxious weed> (possibly extirpated), Lens culinaris Medik. are agricultural weeds. Caragana arborescens Lam. only cult. Ononis alopecuroides L. <Noxious weed>, Sphaerophysa salsula (Pall.) DC. <Noxious weed> all evidently extirpated. Cercidium moved to Parkinsonia; Chamaecytisus to Cytisus; Psoralidium lanceolatum to Ladeania. —Scientific Editors: Martin F. Wojciechowski, Thomas J. Rosatti.

Key to Fabaceae

GLEDITSIA HONEY LOCUST
Tree, generally armed (generally unarmed in cultivation); ± dioecious.
Leaf: of new stems alternate, irregularly odd-2- pinnate, of old stems on spurs, odd-1- pinnate.
Inflorescence: on spurs, ± catkin-like, pendent.
Staminate inflorescence: flowers clustered, pedicels 0 to short.
Pistillate or bisexual inflorescence: flowers generally spaced, pedicels > short.
Flower: radial, perianth hairy, sepals, petals each 3–5, ± alike except petals > sepals; stamens generally 5–7[8].
Fruit: ± indehiscent, oblong to ovate, compressed side-to-side yet plump, pulpy, dry, leathery in age.
Seed: [1]many.
13–16 species: e North America, South America, e&se Asia. (J.G. Gleditsch, German botanist, 1714–1786) [Randall & Meyers-Rice 1997 Madroño 44:399–400; Schnabel & Wendel 2003 Amer J Bot 90:310–320]
Unabridged species in genus: [13–16 species: 2–3 in e North America, 1 in South America, 8–10 in e&se Asia.]
Unabridged references: [Gordon 1966 Ph.D. Dissertation, Indiana Univ; Schnabel & Wendel 2003 Amer J Bot 90:310–320; Randall & Meyers-Rice 1997 Madroño 44:399–400]

G. triacanthos L. HONEY LOCUST
NATURALIZED

Leaf: deciduous; 2- pinnate leaves with 1° leaflets 7–17, 2° leaflets 4–20, elliptic to oblong, 1.3–2.5 cm, glabrous; 1- pinnate leaves with leaflets 20–28, 1.5–3.5 cm.
Staminate inflorescence: 1–several per spur, 3.5–8 cm, simple or branched at tip.
Pistillate inflorescence: 1 per spur, 3–5 cm, simple.
Flower: perianth yellow-green, ± 3 mm ( staminate), 4–5 mm ( pistillate flower); stamens 5–7.
Fruit: 1–3 per peduncle, 20–40 cm, 2.5–3 cm wide, generally curved, often twisted, hairy in youth, brown, glabrous, shiny, persistent in age.
Uncommon. Moist riparian to dry upland woodland; < 800 m. Sacramento Valley, Peninsular Ranges, expected elsewhere; c&e United States, widely cultivated. May–Jun {Weed listed by BAEDN and Cal-IPCI} [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.