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POLEMONIACEAE PHLOX FAMILY

Robert Patterson, family description, key to genera

Annual, perennial herb, shrub, vine. Leaf: simple or compound, cauline (or most basal), alternate or opposite; stipules 0. Inflorescence: cymes, heads, clusters, or flower 1; bracts in involucres or not. Flower: sepals generally 5, fused at base, translucent membrane generally connecting lobes, torn by fruit; corolla generally 5-lobed, radial or bilateral, salverform to bell-shaped, throat often well defined; stamens generally 5, epipetalous, attached at >= 1 level, filaments of >= 1 length, pollen white, yellow, blue, or red; ovary superior, chambers generally 3, style 1, stigmas generally 3. Fruit: capsule. Seed: 1–many, when wetted swelling or not, gelatinous or not.
26 genera, 314 species: America, northern Europe, northern Asia; some cultivated (Cantua, Cobaea (cup-and-saucer vine), Collomia, Gilia, Ipomopsis, Linanthus, Phlox). [Porter & Johnson 2000 Aliso 19:55–91] Leptodactylon moved to Linanthus. —Scientific Editors: Robert Patterson, Thomas J. Rosatti.

Key to Polemoniaceae

GILIA

J. Mark Porter

Annual. Stem: decumbent to erect, glabrous, hairy, glandular, or tufted-woolly-hairy. Leaf: simple, 1–2-pinnate-lobed or -dissected, generally alternate, tips acute, acuminate, or mucronate; basal generally in rosette, entire, toothed, or 1–2-pinnate-lobed; cauline generally reduced. Inflorescence: flowers 1–many in bract axils. Flower: calyx membranous between lobes, membranes splitting or expanding in fruit; corolla > calyx, lobes generally ovate, acute or acuminate. Fruit: spheric to ovoid; chambers 3; valves separating from top, to base and detaching or not to base and staying attached to receptacle. Seed: 3–many, yellow to brown, gelatinous when wet.
± 40 species: western North America, South America. (Filippo L. Gilii, Italian naturalist, 1756–1821) Stamens, styles said to be exserted protrude beyond fused part of corolla, that is, beyond corolla throat. Other taxa in TJM (1993) moved to Aliciella, Lathrocasis, Linanthus, Navarretia, Saltugilia. Gilia mexicana A.D. Grant & V.E. Grant recently found in San Diego Co.
Unabridged note: Gilia can be challenging to identify. Several small-flowered species are only cryptically different from one another and many traits overlap. In order to successfully use the key, it is important to note pattern and distribution of flower color at the time of collection, as it may fade upon drying. Depauperate specimens may be particularly difficult if not impossible to identify using ordinary means. Descriptions of flowers follows previous authors (Day 1993; Grant & Grant 1956) in subdividing fused part of corolla into tube (basal part with parallel sides) and throat (flared part distal to tube). It is important to note that in some cases these definitions do not refer to homologous parts of the corolla.

Key to Gilia

G. brecciarum M.E. Jones BREAK GILIA, NEVADA GILIA
NATIVE
Occasionally with skunk-like odor. Stem: 8–35 cm, densely tufted-woolly-hairy near base; branches spreading, decumbent, or erect. Leaf: basal in suberect cluster, 2–5 cm, 2-pinnate-lobed, lobes ± irregular, lanceolate, > leaf axis width, axis not strap-shaped; cauline axis wide, upper cauline leaf lobes generally finger-like, terminal lobe wider than lateral. Inflorescence: clusters; pedicel glands dense, minute, black, stalked. Flower: calyx 2.5–6 mm, lobes spreading, acute, thick, densely fine-black-glandular, wider than membranes, membranes generally purple-tinged; corolla 8–20 mm; stamens exserted or ± so. Fruit: 4–7 mm, ± = calyx, widely ovoid. Seed: 18–33.
2n=18. Subspecies intergrade ± in southern California. [Online Interchange]

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Next taxon: Gilia brecciarum subsp. brecciarum

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 20 2014
Citation for this treatment: [Author of taxon treatment] 2012. Gilia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=26872, accessed on Apr 20 2014

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click for enlargement Gilia brecciarum
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2005 Christopher L. Christie

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.