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POACEAE (Gramineae) GRASS FAMILY

James P. Smith, Jr., except as noted

Annual to woody perennial herb; roots generally fibrous. Stem: generally round, hollow; nodes swollen, solid. Leaf: alternate, 2-ranked, generally linear, parallel-veined; sheath generally open; ligule membranous or hairy, at blade base. Inflorescence: various (of generally many spikelets). Spikelet: glumes generally 2; florets (lemma, palea, flower) 1–many; lemma generally membranous, sometimes glume-like; palea generally ± transparent, ± enclosed by lemma. Flower: generally bisexual, minute; perianth vestigial; stamens generally 3; stigmas generally 2, generally plumose. Fruit: grain, sometimes achene- or utricle-like.
650–900 genera; ± 10550 species: worldwide; greatest economic importance of any family (wheat, rice, maize, millet, sorghum, sugar cane, forage crops, ornamental, weeds; thatching, weaving, building materials). [Barkworth et al. 2003 FNANM:25; Barkworth et al. 2007 FNANM:24] Generally wind-pollinated. Achnatherum, Ampelodesmos, Hesperostipa, Nassella, Piptatherum, Piptochaetium, Ptilagrostis moved to Stipa; Elytrigia, Leymus, Pascopyrum, Pseudoroegneria, Taeniatherum to Elymus; Hierochloe to Anthoxanthum; Lolium, Vulpia to Festuca; Lycurus to Muhlenbergia; Monanthochloe to Distichlis; Pleuraphis to Hilaria; Rhynchelytrum to Melinis. The following taxa (in genera not included here), recorded in California from historical collections or reported in literature, are extirpated, lacking vouchers, or not considered naturalized: Acrachne racemosa (Roth) Ohwi, Allolepis texana (Vasey) Soderstr. & H.F. Decker, Amphibromus nervosus (Hook. f.) Baill., Axonopus affinis Chase, Axonopus fissifolius (Raddi) Kuhlm., Coix lacryma-jobi L., Cutandia memphitica (Spreng.) K. Richt., Dinebra retroflexa (Vahl) Panz., Eremochloa ciliaris (L.) Merr., Eustachys distichophylla (Lag.) Nees, Gaudinia fragilis (L.) P. Beauv., Miscanthus sinensis Andersson, Neyraudia arundinacea (L.) Henrard, Phyllostachys aurea Rivière & C. Rivière, Phyllostachys bambusoides Siebold & Zuccarini, Rottboellia cochinchinensis (Lour.) Clayton, Schedonnardus paniculatus (Nutt.) Branner & Coville, Schizachyrium cirratum (Hack.) Wooton & Standl., Schizachyrium scoparium (Michx.) Nash, Themeda quadrivalvis (L.) Kuntze, Thysanolaena latifolia (Hornem.) Honda, Tribolium obliterum (Hemsl.) Renvoize, Zea mays L., Zizania palustris L. var. interior (Fassett) Dore, Zoysia japonica Steud. Paspalum pubiflorum E. Fourn., Paspalum quadrifarium Lam., are now reported for southern California (J Bot Res Inst Texas 4:761–770). See Glossary p. 30 for illustrations of general family characteristics. —Scientific Editors: James P. Smith, Jr., J. Travis Columbus, Dieter H. Wilken.
Unabridged references: [Hitchcock 1951 Manual grasses US, USDA Misc Publ 200; Clayton & Renvoise 1986 Kew Bull Add Series 13]

Key to Poaceae

FESTUCA FESCUE, RYE GRASS

James P. Smith, Jr. & Susan G. Aiken

Annual, perennial herb, loosely to densely cespitose, generally ± glabrous, rhizomes present or 0; bisexual, dioecious in Festuca kingii. Stem: decumbent, ascending to erect, to 2 m. Leaf: basal and cauline; sheath generally persisting; collar generally glabrous; ligule generally < 1 mm, membranous, truncate, minutely fringed; blade flat or rolled, claw-like basal auricles generally 0. Inflorescence: panicle-like, branches dense and appressed to open and spreading or spike-like. Spikelet: glumes 2, < lowest floret, unequal, lower sometimes minute, 1–3-veined, upper 3–5-veined; axis breaking above glumes and between florets, florets (1)2–10(17), generally bisexual; lemma 3–5(7)-veined, base generally glabrous, tip entire, occasionally minutely 2-toothed, awn 0 or terminal, generally straight, glabrous; palea ± = lemma; stamens 3 or 1. Fruit: obovoid to elongate, free from or ± adhering to palea and lemma.
500+ species: cosmopolitan. (Latin: straw, stem or straw-like weed) [Darbyshire 2007 FNANM 24:443–448; Darbyshire & Pavlick 2007 FNANM 24:389–443; Lonard 2007 FNANM 24:448–454] As treated here, including ×Festulpia, ×Festulolium, Leucopoa, Lolium, ×Schedolium, ×Schedololium, Schedonorus, and Vulpia. Lolium rigidum Gaudin (combination in Festuca evidently not available), native to Europe and Africa, has been found sporadically as an urban and agricultural weed, but has not been collected in recent years and is doubtfully naturalized.
Unabridged references: [Aiken et al. 1996 onwards @ http://biodiversity.uno.edu/delta/festuca/index.htm [broken 21.vii.2010]; Lonard & Gould 1974 Madroño 22:217–230; Wilson 1999 Ph.D. Dissertation Oregon State Univ]

Key to Festuca

F. kingii (S. Watson) Cassidy
NATIVE
Perennial herb, rhizomes short; dioecious. Stem: 3–8 dm, tufted; nodes visible. Leaf: sheath glabrous to densely hairy, conspicuously persisting, generally red-brown with age; ligule 0.5–3.5 mm, back puberulent; blade 3–30 cm, 1.5–7 mm wide, ± flat, stiffly erect, glaucous. Inflorescence: branches appressed, some with spikelets near base. Staminate spikelet: 5–10(12) mm; lower glume 3–5.5 mm, upper 4–6.5 mm; florets 2–5; lemma 5–8 mm, finely scabrous, awn 0; anthers 3.5–5 mm; pistil reduced or 0. Pistillate spikelet: 5–8 mm; lower glume 3–5 mm, upper 3.5–6 mm; florets 2–5; lemma 5–8 mm, finely scabrous, awn 0; stamens sterile or 0; ovary tip hairy. Fruit: plump, minutely beaked.
2n=56. Dry, sandy places, sagebrush plains to subalpine forest; > 1550 m. s High Sierra Nevada, San Bernardino Mountains, Great Basin Floristic Province; to Oregon, New Brunswick. [Leucopoa kingii (S. Watson) W.A. Weber] Jun–Aug [Online Interchange]
Unabridged synonyms: [Hesperochloa kingii (S. Watson) Rydb.]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 29 2014
Citation for this treatment: [Author of taxon treatment] 2013. Festuca, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=25812, accessed on Aug 29 2014

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Geographic subdivisions indicated for the distribution of Festuca kingii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.