Jepson Herbarium
		University of California, Berkeley

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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |

Annual, perennial herb, or ± woody, generally glandular-hairy.
Leaf: simple to compound, basal and cauline; cauline alternate or opposite, stipules 2, ± on stem.
Inflorescence: cyme or pseudo- umbel or 1–2-flowered.
Flower: bisexual [ unisexual], radial or ± bilateral; sepals 5, free, overlapping in bud; petals generally 5, free, generally with nectar glands at base; stamens generally 5,10[15]; staminodes scale-like or 0; ovary generally 5-lobed, upper part elongating into beak in fruit, chambers 5, placentas axile, style 1, stigmas 5, free, persistent in fruit.
Fruit: septicidal [ loculicidal], mericarps 5, dry, generally 1-seeded, each persistent on 1 of 5 linear segments of beak that separate from central column by curving or coiling upward.
6 genera, ± 750 species: temperate, ± tropics. Some cultivated for ornamental, perfume oils. [Bakker et al. 2006 Taxon 55:887–896] —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Bakker, F. T., Breman, F., & Merckx, V. 2006. DNA sequence evolution in fast-evolving mitochondrial DNA nad1 exons in Geraniaceae and Plantaginaceae. Taxon 55: 887–896; Price & Palmer 1993 Ann Missouri Bot Gard 80:661–671]

Key to Geraniaceae

Annual, perennial herb.
Leaf: simple to pinnately compound, cauline opposite; blade lanceolate to reniform in outline, puberulent or short-hairy, base cordate to truncate.
Inflorescence: umbel.
Flower: radial; stamens 5, free, alternate 5 scale-like staminodes.
Fruit: mericarp body indehiscent, fusiform, 1-seeded, base sharply pointed, top generally with 1 pit on each side of beak segment, pits subtended by 1–4 ridges or not; beak segments stiffly hairy adaxially, generally twisted.
± 74 species: temperate America, Eurasia, n Africa, Australia. (Greek: heron, from bill-like fruit) [Fiz et al. 2006 Syst Bot 31:739–763] Some cultivated for forage, dyes; "beak segments" sometimes called "awns" elsewhere. Erodium macrophyllum moved to genus California.
Unabridged references: [Guittonneau 1972 Boissiera 20:1–154]

Annual.
Stem: decumbent to ascending, 1–5 dm, ± glandular-hairy.
Leaf: compound; lower 3–10 cm, blade > petiole, ovate to oblanceolate in outline, sparsely hairy; leaflets 9–13, deeply dissected, ultimate segments 1–2 mm wide.
Flower: sepals 3–5 mm, tip bristly; petals ± = sepals, pink to purple, base with veins generally darker.
Fruit: body 4–7 mm, pits ± round, glabrous, subtended by 1–2 glabrous ridges; style column 2–5 cm.
2n=20,36,40,42,54,60. Open, disturbed sites, grassland, scrub; < 2000 m. California;

Previous taxon: Erodium brachycarpum
Next taxon: Erodium cygnorum

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.