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PRIMULACEAE PRIMROSE FAMILY

Anita F. Cholewa, except as noted

Annual, perennial herb, or subshrub, glabrous to glandular-hairy. Leaf: simple, ± basal, petioled or not; stipules 0. Inflorescence: scapose umbel, subtended by involucre. Flower: bisexual, radial; parts in 4s or 5s (6s); calyx deeply lobed, often persistent; corolla lobes erect or spreading to reflexed; stamens epipetalous, opposite corolla lobes; ovary superior, 1-chambered, placenta free-central, style 1, stigma head- or dot-like. Fruit: capsule, 2–7-valved or circumscissile. Seed: small, few to many.
± 8 genera, 600 species: northern hemisphere; several ornamental (Primula). [Martins et al. 2003 Plant Syst Evol 237:75–85] Based on molecular evidence, non-rosette terrestrial members of Primulaceae as treated in TJM (1993) moved to Myrsinaceae, and Samolus to Theophrastaceae; based on the same evidence, Primulaceae has been treated alternatively to include all of Myrsinaceae and Theophrastaceae. Taxa of Dodecatheon in TJM2 treated here in Primula. —Scientific Editors: Bruce G. Baldwin, David J. Keil, Thomas J. Rosatti.
Unabridged references: [Källersjö et al. 2000 Amer J Bot 87:1325–1341; Lens et al. 2005 Syst Bot 30:163–183; Martins et al. 2003 Plant Syst Evol 237:75–85; Oberprieler & Hellwig 2003 Plant Syst Evol 237: 75–85; Stähl & Anderberg, in Kubitzki et al. 2004 Fam Generally Vasc Plant 6: 266–281]
Unabridged note: Recent molecular work has led to new understanding of relationships in Ericales. As treated here, non-rosette, terrestrial members of Primulaceae s.l. have been removed from that family and inserted in Myrsinaceae. Myrsinaceae, then, is characterized by synapomorphies of dark dots or streaks on stems, leaves, or flowers, short corolla tubes, seeds immersed in the placenta, and — for woody members — wood lacking rays or with only multiseriate rays. Primulaceae, Myrsinaceae, Theophrastaceae, and Maesaceae (a recent segregate of tropical trees), now constitute a closely related, monophyletic clade. A more recently proposed, alternative taxonomy treats all members of that large clade within an expanded Primulaceae.

Key to Primulaceae

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 19 2014
Citation for this treatment: [Author of taxon treatment] 2013. Primulaceae, Revision 1, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=232, accessed on Sep 19 2014

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