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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
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Annual, perennial herb, often rhizomed or stoloned, often of wet open places; roots fibrous; monoecious, dioecious, or flowers bisexual. Stem: generally 3-sided, generally solid. Leaf: generally 3-ranked; base sheathing, sheath generally closed, ligule generally 0; blade (0 or) linear, parallel-veined. Inflorescence: spikelets generally arranged in head-, spike-, raceme-, or panicle-like inflorescences; flower generally sessile in axil of flower bract, enclosed in a sac-like structure (perigynium) or generally not. Flower: unisexual or bisexual, small, generally wind-pollinated; perianth 0 or generally bristle like; stamens generally 3, anthers attached at base, 4 chambered; ovary superior, chamber 1, ovule 1, style 2–3-branched. Fruit: achene, 2–3 sided.
± 100 genera, 5000 species: especially temperate. [Gilmour et al. 2013 Kew Bull 68:85–105] Difficult; taxa differ in technical characters of inflorescence, fruit. In Carex and Kobresia, what appear to be individual pistillate flowers in fact are highly reduced inflorescences (whether or not the same applies to staminate flowers is still under debate). In some other works (e.g., FNANM) these are called spikelets, and they are treated as being arranged in spikes. Here and in TJM (1993), what appear to be individual pistillate flowers are called pistillate flowers in Carex (and they are treated as being arranged in spikelets), but spikelets in Kobresia (and they are treated as being arranged into spikes). Though internally inconsistent, the approach here is consistent with traditional usage, and reflects a preference for character states that may be determined in the field. Molecular, morphological, and embryological evidence indicates that Eriophorum crinigerum is to be segregated to a new genus, as Calliscirpus criniger (A. Gray) C.N. Gilmour et al., along with a second, newly described species, Calliscirpus brachythrix C.N. Gilmour et al. (Gilmour et al. 2013); key to genera modified by Peter W. Ball to include Calliscirpus. —Scientific Editors: S. Galen Smith, Thomas J. Rosatti, Bruce G. Baldwin.
Unabridged references: [Ball et al. 2002 FNANM 23:1–608; Bruhl 1995 Australian Syst Bot 8:125–305; Tucker 1987 J Arnold Arbor 68:361–445;]
Key to Cyperaceae
Annual, perennial herb, glabrous. Stem: generally > 1, erect, 2–100 cm, 3-sided or cylindric. Leaf: basal; blade 0 or linear. Inflorescence: terminal, generally ± umbel-like, with spikes on rays; inflorescence bracts 1–9, leaf-like, spreading or erect; rays <= 20 cm; spikelets flat to not flat; flower bracts 2-ranked, 2–36, each with 1 flower in axil. Flower: bisexual; perianth 0; stamens (1–2) 3; stigmas 2–3. Fruit: (ob)ovoid, generally 3-angled, brown, generally not beaked.Key to Cyperus
± 600 species: temps, tropics worldwide. (Greek: name for European Cyperus longus) [Tucker 1994 Syst Bot Monogr 43:1–213] Mature fruit generally needed for identification. Cyperus gracilis R. Br., Cyperus owanii Boeck. [Cyperus ligularis L., misappl.], Cyperus regiomontanus Britton, Cyperus retrorsus Chapm., Cyperus virens Michx., urban weeds in California. Cyperus prolifer Lam. not in California.
Perennial herb 1–50 cm; rhizomes short, ± 1 mm thick. Inflorescence: inflorescence bracts 2–3, 1–15 cm; rays 0–2, 0–5(40) mm; spikes dense, head-like, 5–15 mm wide; spikelets 3–25, 3–9 mm, linear-oblong, flat; flower bracts appressed, 3–9, 1.5–2.2 mm, ovate, light brown to black, lateral veins 0. Flower: stigmas 2. Fruit: 1.2–1.4 mm, ovoid to elliptic, 2-sided, brown.
Marshes, swamps, moist roadsides; < 1500 m. Klamath Ranges, North Coast Ranges, Cascade Range, Sierra Nevada Foothills, n High Sierra Nevada, Great Central Valley, Central Western California, South Coast, Peninsular Ranges, East of Sierra Nevada; to Oklahoma; South America, South Africa. [Cyperus diandrus Torr. var. castaneus S. Watson; Cyperus niger var. capitatus (Britton) O'Neill; Cyperus niger var. castaneus (S. Watson) Këk.] Jul–Nov [Online Interchange]
Previous taxon: Cyperus laevigatus
Next taxon: Cyperus odoratus
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 28 2015
Citation for this treatment: [Author of taxon treatment] 2013. Cyperus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=21881, accessed on Nov 28 2015
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|Geographic subdivisions indicated for the distribution of Cyperus niger|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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