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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Annual, perennial herb, subshrub, generally twining or trailing. Leaf: 0 or alternate. Inflorescence: cyme or flowers 1 in axils; bracts subtending flowers 0 or 2. Flower: bisexual, radial; sepals (4)5, ± free, overlapping, persistent, often unequal; corolla generally showy, generally bell-shaped, ± shallowly 5-lobed, generally pleated and twisted in bud; stamens 5, epipetalous; pistil 1, ovary superior, chambers generally 2, each generally 2-ovuled, styles 1–2. Fruit: generally capsule. Seed: 1–4(6).
55–60 genera, 1600–1700 species: warm temperate to tropics; some cultivated for food or as ornamental (Ipomoea). [Stefanovic et al. 2003 Syst Bot 28:791–806] Monophyletic only if Cuscutaceae included, as treated here. Ipomoea cairica (L.) Sweet, Ipomoea hederacea Jacq. [Ipomoea nil L., misappl.], Ipomoea indica (Burm.) Merr. (including Ipomoea mutabilis Ker Gawl.), Ipomoea purpurea (L.) Roth, Ipomoea triloba L., all included in TJM (1993), not naturalized. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Stefanovic et al. 2002 Amer J Bot 89:1510–1522]
Key to Convolvulaceae
Vine, annual (perennial herb if on perennial host), not in contact with ground, attached to, holoparasitic on host by many small, specialized roots (haustoria) along stem, generally glabrous. Stem: thread-like, ± green, yellow, orange, or ± red. Leaf: 0 or scale-like, alternate, ± 2 mm. Inflorescence: generally cyme, head- to panicle-like (flowers 1), subtended by 0–3 bracts. Flower: bisexual, radial, parts generally in 4s or 5s; calyx generally divided 2/5–3/5, persistent generally ± cream-white; corolla generally ± white, persistent (withered in fruit) or not, tube cup-shaped to cylindric, bulged or horizontally ridged below lobes or generally not, generally with scales subtending stamens, lobes alternate stamens, erect to reflexed; ovary superior, chambers 2, each 2-ovuled, styles 2, generally free, persistent, stigmas 2, generally spheric, persistent. Fruit: capsule, generally indehiscent to irregularly dehiscent (or circumscissile near base), spheric to ovoid, depressed or not, thickened and/or raised around generally inconspicuous opening between styles or not. Seed: 1–4; coat papillate when hydrated, honeycombed when dry, (rarely neither, with cells ± rectangular, in ± jigsaw-puzzle-like arrangement); embryo generally slender, 1–3-coiled.Key to Cuscuta
± 180 species: cosmopolitan, especially warmer regions of western hemisphere and Polynesia. (Aramaic, Hebrew; to cover, from habit) [Costea & Stefanovic 2009 Syst Bot 34:570–579] By persistent, withered corolla, fruit may be "capped" (corolla on top of fruit), "surrounded" (fruit at least in part visible, corolla ± loosely around fruit), or "enclosed" (fruit not visible, corolla ± tightly around fruit). Cuscuta pentagona Engelm. excluded.
Unabridged etymology: (Aramaic, Hebrew; from the verb K-S-Y (Kaph, Shin, Yodh), to cover, from habit)
Unabridged references: [Costea et al. 2005 Brittonia 57:264–272; Costea et al. 2006 Sida 22:151–175, 177–195, 197–207, 209–225; Costea & Stefanovic 2009. Cuscuta jepsonii (Convolvulaceae), an invasive weed or an extinct endemic? Amer J Bot 96:1744–1750; Costea et al. 2009. Untangling the systematics of salt marsh dodders: Cuscuta pacifica a new segregate species from Cuscuta salina (Convolvulaceae). Syst Bot 34:787–795; Costea & Stefanovic. 2009. Molecular phylogeny of Cuscuta californica complex (Convolvulaceae) and a new species from New Mexico and Trans-Pecos. Syst Bot 34:570–579; Costea & Tardif 2006 Canad J Plant Sci 86:293–316]
Inflorescence: head- to raceme-like, flowers 6–25(30); pedicels 0.3–2.5(3.5) mm. Flower: (1.9)2.1–3.6 mm, membranous, not papillate, parts generally in 5s; calyx generally ± = corolla tube, cup-shaped, divided 2/5–3/5, lobes ovate-triangular, veined, shiny, bases overlapped, margins entire, tips obtuse to rounded; corolla tube (1.1)1.5–1.9 mm, bell-shaped, scales >= corolla tube, oblong-ovate to spoon-shaped, rounded, uniformly densely fringed, lobes spreading, triangular-lanceolate, ± = tube, margins entire, tips acute to acuminate, incurved; filaments 0.4–0.7 mm, anthers exserted, (0.3)0.4–0.5 mm, widely elliptic; styles 0.8–1.6 mm, generally = ovary. Fruit: 1.3–2.8 mm, 1.9–3.8 mm wide, spheric-depressed to depressed, not thickened or raised around opening between styles, translucent or not, corolla enveloping 1/3 or less of fruit. Seed: 4, 1.1–1.5 mm, 0.9–1.1 mm wide, ± spheric to widely elliptic.
2n=56. Generally on herbs, many crops, roadsides, fields; < 500 m. North Coast, Inner North Coast Ranges, Sierra Nevada Foothills, Great Central Valley, Central Coast, South Coast; worldwide. Treated as synonym of Cuscuta pentagona Engelm. in TJM (1993), but more recent work has shown that Cuscuta campestris and Cuscuta pentagona are distinct species, and that the latter does not occur in California. May–Nov [Online Interchange]
Unabridged synonyms: [Cuscuta pentagona var. calycina Engelm.]
Previous taxon: Cuscuta californica var. papillosa
Next taxon: Cuscuta cephalanthi
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Cuscuta, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=21462, accessed on Oct 1 2014
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|Geographic subdivisions indicated for the distribution of Cuscuta campestris|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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