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CUCURBITACEAE GOURD FAMILY

John M. Miller & Robert L. Schlising

Annual, perennial herb; hairs often hard from calcium deposits; generally monoecious. Stem: trailing or climbing, 1–many; tendril generally 1 per node, often branched. Leaf: generally simple, alternate, generally palmate-lobed and -veined, petioled; stipules 0. Inflorescence: at nodes; staminate flowers in racemes, panicles, small clusters, (or 1); pistillate flowers generally 1. Flower: unisexual [bisexual], radial; hypanthium > ovary; calyx generally 5-lobed (or ± 0); corolla rotate to cup- or bell-shaped, generally 5-lobed; stamens 3–5 (or ± 1–3 from fusion), anthers twisted together, often > filaments; ovary ± inferior, chambers 3–5, placentas parietal, styles 1–3, stigmas generally lobed, large. Fruit: berry, drying or not, or capsule, irregularly dehiscent, often gourd- or melon-like. Seed: 1–many.
100 genera, 700 species: especially tropics; some cultivated (Citrullus, watermelon; Cucumis, cucumber; Cucurbita, gourd, pumpkin, squash; Luffa, loofah; Sechium, chayote). [Schaefer et al. 2009 Proc Roy Soc London Ser B, Biol Sci 276:843–851] Several cultivated species included Bryonia dioica reported as waifs in California (Howell 1958 Wasmann J Biol 16:1–157), but none recently. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Jeffrey 1990 An outline classification of the Cucurbitaceae, In: Bates, Robinson, and Jeffrey (eds.), Biology and Utilization of the Cucurbitaceae 17:449–463, Cornell Univ Press, Ithaca, NY; Kocyan et al. 2007 Molec Phylogen Evol 44:553–577; Lira, Rodriguez-Jimenez, Alvarado, Rodriguez, Castrejon, Dominguez-Mariani 1998 Acta Bot Mex 42: 43–77; McVaugh 2001 Cucurbitaceae In W.R. Anderson (ed), Flora Novo-Galiciana 3:483–652, Univ Michigan Herbarium, Ann Arbor, Michigan; Schaefer et al. 2008 Syst Bot 33:125–132]

Key to Cucurbitaceae

CUCURBITA GOURD, FIELD PUMPKIN, SQUASH
Annual, perennial herb, from large, fleshy, tuber-like root. Stem: glabrous to scabrous; tendril branched or not. Leaf: lanceolate to round, entire to deeply lobed. Inflorescence: flowers 1 per node, staminate, pistillate at different nodes. Flower: corolla > 2 cm wide (staminate generally < pistillate), deeply cup-shaped, yellow to orange, fused part 4–12 cm, lobes generally recurved; stigmas 3, 2-lobed. Fruit: spheric to oblong, indehiscent; rind firm, smooth to rough or grooved. Seed: many, < 20 mm, ± ovate, ± flat; margin thick or raised.
12–14 species: America. (Latin: gourd) [Sanjur et al. 2002 Proc Natl Acad Sci USA 99:535–540]
Unabridged references: [Montes-Hernandez & Eguiarte 2002 Amer J Bot 89:1156–1163; McVaugh 2001 Cucurbita In W.R. Anderson (ed), Flora Novo-Galiciana 3:510–529, Univ Michigan Herbarium, Ann Arbor, Michigan; Decker 1988 Econ Bot 42:4–15]

Key to Cucurbita

C. digitata A. Gray FINGER-LEAVED GOURD
NATIVE
Herbage ± scabrous; tendril branched ± from base. Leaf: 3–9 cm, palmately lobed, green, main veins lighter, lobes generally 5, free ± to petiole, ± lance-linear. Flower: corolla 3–5 cm, yellow. Fruit: 7–8 cm wide, spheric to oblong, dark green, ± mottled, with several narrow, ± white stripes. Seed: 10–11 mm, white.
2n=40. Uncommon. Sandy, open or shrubby places; < 1200 m. South Coast, Peninsular Ranges, Sonoran Desert; to New Mexico, northern Mexico. Aug–Oct [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Cucurbita, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=21360, accessed on Oct 22 2014

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Geographic subdivisions indicated for the distribution of Cucurbita digitata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.