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CRASSULACEAE STONECROP FAMILY

Steve Boyd, except as noted

Annual to shrub [(± tree-like or climbing)], fleshy. Leaf: generally simple, alternate or opposite, in dense to open, basal (or terminal) rosettes or basal and cauline, not in rosettes, reduced distally or not, margin often ± red. Inflorescence: generally cyme, generally bracted. Flower: generally bisexual; sepals generally 3–5, generally ± free; petals generally 3–5, ± free or fused; stamens >> to = sepals, epipetalous or not; pistils generally 3–5, simple, fused at base or not, ovary 1-chambered, placenta 1, parietal, ovules 1–many, style 1. Fruit: follicles, generally 3–5. Seed: 1–many, small.
± 33 genera, ± 1400 species: ± worldwide, especially dry temperate; many cultivated for ornamental. [Eggli (ed.) 2003 Illus Handbook Succulent Plants 6 (Crassulaceae). Springer] Water-stressed plants often ± brown or ± red. Consistent terminology regarding leaves, bracts difficult; in taxa with rosettes (e.g., Aeonium, Dudleya, some Sedum), structures in rosettes are leaves, those on peduncles are bracts, and those subtending flowers are flower bracts; in taxa where inflorescence is terminal, rosette leaves may "become" bracts as stem rapidly elongates to form inflorescence. Seed numbers given per follicle. —Scientific Editor: Thomas J. Rosatti.

Key to Crassulaceae

CRASSULA
Annual, perennial herb, shrub, glabrous (hairy). Stem: erect to decumbent, branched or not. Leaf: opposite, 0.1–7 cm, linear to deltate or obovate, bases fused, ± sheathing; margins generally entire. Inflorescence: terminal panicle or flowers 1 in axils of leaves, either 2 per node, axillary, or 1 per node, terminal but appearing axillary by overtopping of main axis. Flower: erect, sepals 3–5, ± fused at base; petals 3–5, spreading or recurved, free or ± fused at base; stamens = sepals in number; pistils 3–5. Fruit: spreading to erect. Seed: 0.2–0.6 mm, elliptic to elliptic-oblong (spheric, reniform), generally with longitudinal lines, sometimes ± smooth or papillate, red-brown.
x=(7)8.
± 250 species: especially Africa, annual ± worldwide. (Latin: diminutive of thick) Crassula argentea Thunb., a synonym of Crassula ovata (Mill.) Druce, a waif.
Unabridged references: [Moran 1992 Cactus and Succulent Journal 64:223–231]

Key to Crassula

C. connata (Ruiz & Pav.) A. Berger PYGMY-WEED
NATIVE
Annual, terrestrial. Stem: erect, 2–6(10) cm, branched or not, not rooting at nodes, red in age. Leaf: 1–3(6) mm, ovate to oblong; tip acute to rounded, without awn or point. Inflorescence: flowers (1)2 per node; pedicel 0.2–6 mm. Flower: parts generally in 4s; sepals 0.5–2 mm, lanceolate, acute to acuminate; petals generally < sepals, 0.6–1.2(1.5) mm, narrow-deltate. Fruit: ascending, ovoid. Seed: 1–2, elliptic, shiny, with ± wavy longitudinal lines at 20×.
2n=16. Open areas; < 1500 m. Northwestern California, Cascade Range Foothills, Sierra Nevada Foothills, Great Central Valley, Central Western California, Southwestern California, w Mojave Desert, Sonoran Desert; to Washington, southern British Columbia, Texas, northern Central America; also western South America. Locally abundant. Feb–May [Online Interchange]
Unabridged synonyms: [Crassula connata var. connata; Crassula connata var. erectoides M. Bywater & Wickens; Crassula connata var. eremica (Jeps.) M. Bywater & Wickens; Crassula connata var. subsimplex (S. Watson) M. Bywater & Wickens]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 30 2014
Citation for this treatment: [Author of taxon treatment] 2013. Crassula, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=20582, accessed on Sep 30 2014

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click for enlargement Crassula connata
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© 2003 Steve Matson

Geographic subdivisions indicated for the distribution of Crassula connata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.