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CRASSULACEAE STONECROP FAMILY

Steve Boyd, except as noted

Annual to shrub [(± tree-like or climbing)], fleshy. Leaf: generally simple, alternate or opposite, in dense to open, basal (or terminal) rosettes or basal and cauline, not in rosettes, reduced distally or not, margin often ± red. Inflorescence: generally cyme, generally bracted. Flower: generally bisexual; sepals generally 3–5, generally ± free; petals generally 3–5, ± free or fused; stamens >> to = sepals, epipetalous or not; pistils generally 3–5, simple, fused at base or not, ovary 1-chambered, placenta 1, parietal, ovules 1–many, style 1. Fruit: follicles, generally 3–5. Seed: 1–many, small.
± 33 genera, ± 1400 species: ± worldwide, especially dry temperate; many cultivated for ornamental. [Eggli (ed.) 2003 Illus Handbook Succulent Plants 6 (Crassulaceae). Springer] Water-stressed plants often ± brown or ± red. Consistent terminology regarding leaves, bracts difficult; in taxa with rosettes (e.g., Aeonium, Dudleya, some Sedum), structures in rosettes are leaves, those on peduncles are bracts, and those subtending flowers are flower bracts; in taxa where inflorescence is terminal, rosette leaves may "become" bracts as stem rapidly elongates to form inflorescence. Seed numbers given per follicle. —Scientific Editor: Thomas J. Rosatti.

Key to Crassulaceae

CRASSULA
Annual, perennial herb, shrub, glabrous (hairy). Stem: erect to decumbent, branched or not. Leaf: opposite, 0.1–7 cm, linear to deltate or obovate, bases fused, ± sheathing; margins generally entire. Inflorescence: terminal panicle or flowers 1 in axils of leaves, either 2 per node, axillary, or 1 per node, terminal but appearing axillary by overtopping of main axis. Flower: erect, sepals 3–5, ± fused at base; petals 3–5, spreading or recurved, free or ± fused at base; stamens = sepals in number; pistils 3–5. Fruit: spreading to erect. Seed: 0.2–0.6 mm, elliptic to elliptic-oblong (spheric, reniform), generally with longitudinal lines, sometimes ± smooth or papillate, red-brown.
x=(7)8.
± 250 species: especially Africa, annual ± worldwide. (Latin: diminutive of thick) Crassula argentea Thunb., a synonym of Crassula ovata (Mill.) Druce, a waif.
Unabridged references: [Moran 1992 Cactus and Succulent Journal 64:223–231]

Key to Crassula

C. aquatica (L.) Schönl.
NATIVE
Annual, generally aquatic or on wet substrates. Stem: decumbent, ± erect if stranded, generally branched at base, rooting at nodes. Leaf: 2–6 mm, oblanceolate to linear, tip acute. Inflorescence: flower 1 per node, terminal; pedicel 0.5–20 mm. Flower: parts in 4s; sepals ± 0.5–1.5 mm, ovate to oblong, rounded to obtuse; petals > sepals, 1–2 mm, ovate to oblong. Fruit: erect, oblong, tip subtruncate; suture straight, abruptly outcurved in distal 1/4. Seed: 6–17, elliptic-oblong, ± dull or shiny but not glistening as if wet, with ± regular, continuous, longitudinal lines at 20×.
2n=42. Salt marshes, vernal pools, margins of lakes, ponds; < 3000 m. North Coast, Cascade Range Foothills, Sierra Nevada Foothills, Great Central Valley, Central Western California, Southwestern California (except n Channel Islands); North America, Mexico, northern Eurasia. Mar–Jun(Aug) [Online Interchange]
Unabridged synonyms: [Crassula saginoides (Maxim.) M. Bywater & Wickens]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 17 2014
Citation for this treatment: [Author of taxon treatment] 2012. Crassula, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=20581, accessed on Apr 17 2014

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click for enlargement Crassula aquatica
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© 2013 Anna Bennett

Bioregions in which Crassula aquatica occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.