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Daniel Potter & Barbara Ertter, family description, key to genera; treatment of genera by Daniel Potter, except as noted

Annual to tree, glandular or not. Leaf: simple to palmately or pinnately compound, generally alternate; stipules free to fused (0), persistent to deciduous. Inflorescence: cyme, raceme, panicle, cluster, or flowers 1; bractlets on pedicel ("pedicel bractlets") generally 0–3(many), subtended by bract or generally not. Flower: generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, subtending bractlets ("hypanthium bractlets") 0–5, alternate sepals; sepals generally 5; petals generally 5, free; stamens (0,1)5–many, anther pollen sacs generally 2; pistils (0)1–many, simple or compound, ovary superior to inferior, styles 1–5. Fruit: 1–many per flower, achene (fleshy-coated or not), follicle, drupe, or pome with generally papery core, occasionally drupe-like with 1–5 stones. Seed: generally 1–5 (per fruit, not per flower).
110 genera, ± 3000 species: worldwide, especially temperate; many cultivated for ornamental, fruit, especially Cotoneaster, Fragaria, Malus, Prunus, Pyracantha, Rosa, Rubus. [Potter et al. 2007 Plant Syst Evol 266:5–43] Number of teeth is per leaf or leaflet, not per side of leaf or leaflet, except in Drymocallis. —Scientific Editors: Daniel Potter, Thomas J. Rosatti.
Unabridged references: [Robertson 1974 J Arnold Arbor 55:303–332, 344–401, 611–662]

Key to Rosaceae


Barbara Ertter

1 sp. (Greek: fruit of Arbutus unedo) [Ertter 2007 J Bot Res Inst Texas 1:31–46]

Perennial herb, nonglandular, openly matted. Stem: non-flowering portion creeping, often floating and/or rooting at nodes; flower-stem erect to ascending, 20–50 cm, ± glabrous to ± strigose. Leaf: alternate, odd-1-pinnate to subpalmate, 5–20 cm; leaflets 2–3 per side, toothed, terminal generally ± = lateral, 15–70 mm, oblanceolate-elliptic, distal 1/3 toothed ± 1/4 to midvein, ± glabrous to sparsely strigose. Inflorescence: cyme, ± open, generally < 10-flowered; pedicels straight. Flower: hypanthium ± shallow, 5–10 mm wide; bractlets 5; sepals ± triangular; petals << sepals, 2–6 mm, ± ovate to elliptic, dark red; stamens 20–25, filaments 1.5–2.5 mm, anthers 1–1.5 mm, pollen sac 1, horseshoe-shaped; pistils many, ovaries superior, styles 0.8–2 mm, slender, jointed laterally to fruit. Fruit: achene, 1–1.5 mm, glabrous, glossy, ± brown, sometimes remaining attached to spongy receptacle.
2n=28,42–64. Bogs, marshes; < 2400 m. North Coast, High Cascade Range, n High Sierra Nevada, Sacramento Valley, Warner Mountains; circumboreal. [Potentilla palustris (L.) Scop.] May–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 24 2015
Citation for this treatment: [Author of taxon treatment] 2013. Comarum, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 24 2015

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click for enlargement Comarum palustre
See CalPhotos for additional images
1998 Gary A. Monroe

Geographic subdivisions indicated for the distribution of Comarum palustre Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.