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RHAMNACEAE BUCKTHORN FAMILY

John O. Sawyer, Jr., except as noted

[Perennial herb] shrub, tree, generally erect, often thorny. Leaf: simple, generally alternate, often clustered on short-shoots; stipules generally present, occasionally modified into spines; generally petioled; blade pinnate-veined or 1–5-ribbed from base. Inflorescence: cyme, panicle, umbel, or flowers 1 or clustered in axils or on short-shoots. Flower: generally bisexual, radial; hypanthium subtending, surrounding, or partly fused to ovary; sepals 4–5; petals 0, 4–5, generally clawed; stamens 0, 4–5, alternate sepals, attached to hypanthium top, each generally fitting into a petal concavity; disk (0 or) between stamens, ovary, thin to fleshy, entire or lobed, free from ovary, adherent or fused to hypanthium; ovary superior or ± inferior, chambers [1]2–4, 1–2-ovuled, style 1, stigma entire or 2–3-lobed. Fruit: capsule, drupe.
50–52 genera, 950 species: especially tropics, subtropics some cultivated (Ceanothus; Frangula; Rhamnus; Ziziphus). [Richardson et al. 2000 Amer J Bot 87:1309–1324] —Scientific Editors: Steve Boyd, Thomas J. Rosatti.
Unabridged references: [Brizicky 1965 J Arnold Arbor 45:439–463; Richardson et al. 2000 Kew Bull 55:311–341]

Key to Rhamnaceae

COLUBRINA
Shrub. Stem: branches dense, alternate, rigid, ± white; twigs spreading, not jointed at base, thorn-tipped [or not], generally hairy. Leaf: clustered on short-shoots or not, deciduous [or not]; stipules deciduous; blade entire, 3–5-ribbed from base, generally with round, marginal glands. Inflorescence: umbel-like, few-flowered. Flower: hypanthium hemispheric, adhering to developing fruit; sepals 5; petals 5, = sepals, oblanceolate; stamens 5; disk fleshy, adhering to, ± filling hypanthium; ovary chambers 3, 1-ovuled, stigma 3-lobed. Fruit: capsule, shallow-3-valved, explosively dehiscent (drupe-like, indehiscent).
31 species: warm places worldwide. (Latin: from French for serpent tree) [Bastos 1990 Pesquisas Botanica 41:99–122]
Unabridged references: [Johnston 1971 Brittonia 23:2–53]

C. californica I.M. Johnst. LAS ANIMAS COLUBRINA
NATIVE
Plant < 3 m. Stem: straight, tomentose. Leaf: blade 12–35 mm, oblong to obovate, dull gray-green, hairs silky, denser abaxially, base rounded or wedge-shaped, tip rounded to ± notched, mucronate or not. Inflorescence: 5–10 mm, 3–12-flowered, dense; pedicel 1–2 mm, 2–4 mm in fruit. Flower: appearing after rain; hypanthium ± 3 mm wide. Fruit: 8–10 mm, persisting 3–6 months.
Desert scrub; 240–920 m. s Mojave Desert, Sonoran Desert; Arizona, Mexico. Apr–May [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Colubrina, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=20048, accessed on Sep 1 2014

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Geographic subdivisions indicated for the distribution of Colubrina californica Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.