Jepson Herbarium
		University of California, Berkeley

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Annual to shrub, some aquatic.
Leaf: basal or cauline, alternate or opposite ( whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0.
Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract.
Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, [1]2–4-chambered, style 1, stigma lobes 0 or 2.
Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic CA genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). CA Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.

Key to Plantaginaceae

Annual, often glandular, sometimes brown-staining.
Leaf: opposite; proximal petioled.
Inflorescence: bracted, often interrupted; flowers 1–many in leaf axils.
Flower: calyx lobes 5, generally glabrous on inner surface; corolla ± pea-like, uniformly pale, or generally with pale regions, especially throat and base of upper lip (± uniformly dark in Collinsia greenei), generally glabrous outside, tube short, throat ± angled to tube, ± pouched on upper side, lips generally ± = throat, upper lobes 2, ± reflexed, lower lobes 3, lateral spreading, central lobe keeled, enclosing stamens and style; stamens 4, attached unequally near throat base, spur at base of upper filaments > 1 mm, vestigial, or 0; staminode gland-like; style > 2 mm, stigma minutely 2-lobed.
Fruit: septicidal and loculicidal (valves 2-lobed).
Seed: generally few, ± oblong, generally plump; inner surface ± concave.
± 20 species: North America, especially CA. (Zaccheus Collins, Philadelphia botanist, 1764–1831) Late-season flowers generally atypically small.

Plant 10–40 cm.
Leaf: generally > 6 × longer than wide, ± linear, margins rolled under.
Inflorescence: open, finely scaly and spreading- glandular-hairy; bracts > 2 mm; pedicels generally 1–3(5) per node, > calyx, ascending.
Flower: calyx ± = fruit, lobe tips blunt; corolla generally 8–12(15) mm, >> 2 × calyx, white to blue-purple, throat strongly angled to tube, as wide as long, pouch ± square, prominent, upper lip with projections bulging ± 0.5 mm away from throat opening at base of each lobe; upper filaments hairy at base, basal spur 0–0.5 mm.
Seed: 3–4(+), ± plump, generally unwinged.
n=7. Open conifer forest; 200–2000 m. Klamath Ranges, n Outer North Coast Ranges;

Previous taxon: Collinsia heterophylla var. heterophylla
Next taxon: Collinsia multicolor

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.