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CLEOMACEAE SPIDERFLOWER FAMILY

Robert E. Preston & Staria S. Vanderpool

Annual, perennial herb, shrub, often ill-smelling.
Leaf: generally 1- palmate, generally alternate, generally petioled; stipules generally minute, often bristle-like or hairy; leaflets 0 or 3–7.
Inflorescence: raceme, head, or flowers 1, expanded in fruit; bracts generally 3-parted below, simple above, or 0.
Flower: generally bisexual, radial to ± bilateral; sepals generally 4, free or fused, generally persistent; petals generally 4, free, ± clawed; stamens generally 6, free, exserted, anthers generally coiling at dehiscence; ovary superior, generally on stalk-like receptacle, chamber generally 1, placentas generally 2, parietal, style 1, persistent, stigma generally minute, ± head-like.
Fruit: 2 nutlets or generally capsule, septicidal; valves generally 2, deciduous, leaving septum (frame-like placentas) behind; pedicel generally ± reflexed to spreading.
17 genera, ± 150 species: widespread tropics to arid temperate. [Iltis & Cochrane 2007 Novon 17:447–451] Treated as Capparaceae in TJM (1993). —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Ernst 1963 J Arnold Arbor 44:81–93]

Key to Cleomaceae

CLEOMELLA
Annual, generally glabrous.
Stem: generally ascending to erect, sometimes prostrate when older, generally branched from base, often red-tinged.
Leaf: generally many; petiole generally 7–20 mm; leaflets generally 3.
Inflorescence: raceme, ± terminal, flowers 1 in leaf axils, or both; pedicel generally 4–25 mm.
Flower: parts generally yellow; sepals fused in basal 1/3, generally entire; petals ± sessile, upper 2 often recurved.
Fruit: 2–6 mm, often wider than long; septum elliptic to round; receptacle stalk-like.
Seed: < 10.
± 10 species: arid w North America. (Diminutive of Cleome) [Holmgren 2004 Brittonia 56:103–106]

Key to Cleomella

C. plocasperma S. Watson
NATIVE

Stem: branched generally from upper nodes, 10–55(80) cm, smooth.
Leaf: leaflets 15–45 mm, linear- elliptic.
Inflorescence: raceme, 1–20 cm.
Flower: sepals 0.9–2.2 mm, lanceolate; petals 3.5–7 mm; stamens 8–12 mm, anthers 1.5–1.9 mm; style 0.8–1.2 mm.
Fruit: 4–5 mm; valves ± hemispheric to horn-shaped; receptacle 6–10 mm, spreading to ascending.
Wet, alkaline meadows, greasewood flats, near thermal springs; 875–1710 m. ne San Bernardino Mountains, Great Basin Floristic Province (except Warner Mountains, White and Inyo Mountains), Mojave Desert (except Desert Mountains); to Oregon, Idaho, Utah. May–Oct [Online Interchange]
Unabridged synonyms: [Cleomella plocasperma var. mojavensis (Payson) Crum ex Jeps.; Cleomella plocasperma var. stricta Crum ex Jeps.]

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.