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MONTIACEAE MINER'S LETTUCE FAMILY

John M. Miller, except as noted

Annual to perennial herb; generally fleshy. Stem: 1–many, generally glabrous. Leaf: simple, alternate or opposite. Inflorescence: axillary or terminal; cyme, raceme, panicle, umbel, or flower 1. Flower: bisexual, radial; sepals generally 2(9), free; petals (1)2–19, free or ± fused; stamens 1–many, epipetalous or not, anthers pink, rose, or yellow; ovary superior, chamber 1, ovules 1–many, placenta basal or free-central; styles (0)1–8, generally fused at base, branched. Fruit: capsule, circumscissile or 2–3-valved. Seed: 1–many, shiny or ± pebbly or sculptured, black or gray, generally with oil-filled appendage as food for ants.
± 22 genera, ± 230 species: generally temperate America, Asia, Australia, Europe, Kerguelen Is, New Zealand, southern Africa, poorly represented in Europe; some cultivated (Lewisia, Calandrinia). [Ogburn & Edwards 2009 Amer J Bot 96:391–408] Details of flowers, seeds require 20× magnification. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Applequist et al. 2006 Syst Bot 31:310–319; Nyffeler et al. 2008 Haseltonia 14: 26–36]
Unabridged note: May include Hectorellaceae.

Key to Montiaceae

CISTANTHE

John M. Miller & C. Matt Guilliams

Annual [perennial herb], ± fleshy, from taproot or fibrous roots, generally glabrous. Stem: generally several, generally spreading to ascending. Leaf: cauline [basal and cauline], simple, linear to spoon-shaped, fleshy; basal rosetted. Inflorescence: scapose, raceme, panicle, or umbel, bracts leaf-like or not; bracts generally < sepals, scarious; flowers generally on 1 side of axis, deciduous or not in fruit. Flower: sepals 2, ovate, green, scarious-margined or not, persistent in fruit; petals 5, > sepals; stamens 5–10, anthers yellow; style 1, stigmas 3. Fruit: 3-valved, ovoid. Seed: 6–many, dull to shiny, black.
25–35 species: America. (Greek: rockrose-flower) [Hershkovitz 2006 Gayana Bot 63:13–74]
Unabridged references: [Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz 1991a Ann Missouri Bot Gard 78:1009–1021; Hershkovitz 1991b Phytologia 70:209–225; Hershkovitz 1992 Syst Bot 17:220–238; Hershkovitz & Zimmer 2000 Molec Phylogen Evol 15:419–439; Hinton 1975 Brittonia 27:197–208; Kelley et al. 2003 FNANM 4:460–464.]

Key to Cistanthe

C. ambigua (S. Watson) Hershk. DESERT CISTANTHE
NATIVE
Stem: spreading to erect. Leaf: 1.5–6 cm, linear to ± spoon-shaped, ± cylindric. Inflorescence: panicle of umbel-like clusters, compact; bracts scarious; pedicel 1–3 mm, straight in fruit. Flower: sepals 2–5 mm, glabrous, margins white-scarious; petals 3–5, 2–5 mm; stamens 5–7(10). Fruit: < calyx. Seed: 6–15, 1–2 mm wide, ± elliptic to ovate, smooth, shiny, black.
Desert scrub, sandy to silty soil, often alkaline; < 1000 m. Desert; southwestern Arizona, northwestern Mexico. [Calandrinia ambigua (S. Watson) Howell] Feb–May [Online Interchange]
Unabridged note: Belongs in neither Calandrinia nor Cistanthe, and might better be treated as a separate, monotypic genus, according to Hershovitz (pers. comm.). Expanded author citation: Cistanthe ambigua (S. Watson) Carolin ex Hershk.

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 18 2014
Citation for this treatment: [Author of taxon treatment] 2012. Cistanthe, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=19501, accessed on Apr 18 2014

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click for enlargement Cistanthe ambigua
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Charles Webber 1998 California Academy of Sciences

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.