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POLYGONACEAE BUCKWHEAT FAMILY

Mihai Costea & James L. Reveal, family description, key to genera; treatment of genera revised in consultation with James L. Reveal, except as noted

Annual to shrub [tree]. Stem: nodes swollen or not. Leaf: simple, basal or cauline, generally alternate; ocreae present or 0, generally scarious, persistent or not. Inflorescence: flower clusters in axillary to terminal cyme-, panicle-, raceme-, spike-, umbel- or head-like arrangements, entire inflorescence or main inflorescence branches generally subtended by bracts ("inflorescence bracts"); peduncles present or 0; flower clusters in Eriogoneae-Eriogonoideae subtended by involucre of >= 1 free or ± fused, sometimes awn-tipped bracts ("involucre bracts") or, in Polygonoideae and rarely in Eriogonoideae, not (if bracts completely fused, involucre "tubular"); pedicels in Eriogoneae each often subtended by 2 free, transparent, linear bractlets or in Polygonoideae all subtended by 2+ fused, membranous, wide bractlets. Flower: generally bisexual, small, 1–200 per node; perianth parts 2–6, generally in 2 whorls, free or basally fused, generally petal-like, often ± concave adaxially, often darker at midvein, often turning ± red or ± brown in age; stamens [1]3 or 6–9 in 2 whorls; ovary superior, 1-chambered, ovule 1, styles 1–3. Fruit: achenes, included in or exserted from perianth, generally 3-angled, ovoid or elliptic, generally glabrous.
48 genera, ± 1200 species: worldwide, especially northern temperate; some cultivated for food (Coccoloba, sea-grape; Fagopyrum, Rheum, Rumex) or ornamental (Antigonon, lovechain; Coccoloba; Muehlenbeckia; Persicaria; Polygonum), a few timbered (Coccoloba; Triplaris). Several (Emex; Fallopia; Persicaria; Polygonum; Rumex) are weeds. [Freeman & Reveal 2005 FNANM 5:216–601] Treatment of genera in Eriogonoideae based on monographic work of James L. Reveal. Involucre number throughout is number (1–many) per ultimate grouping, at tips of ultimate branches; flower number is per flower cluster or involucre, unless otherwise stated. Fagopyrum esculentum Moench not naturalized, considered an historical waif (or garden weed ± presently), therefore not treated. —Scientific Editor: Thomas J. Rosatti.

Key to Polygonaceae

CHORIZANTHE SPINEFLOWER
Annual [perennial herb], prostrate to erect, generally hairy. Leaf: basal or cauline, alternate, linear to narrow, awns 0; ocreae 0. Inflorescence: terminal, generally cyme- or head-like, hairy [(glabrous)]; peduncles 0; involucres generally 1–6, tubular, 3–6-ribbed, teeth 3, 5, or 6, awned. Flower: 1(2), glabrous or hairy; perianth white, yellow, red, maroon, or purple, lobes (5)6, entire, notched or lobed to fringed; stamens generally 3, 6, or 9. Fruit: generally brown or black, generally elliptic, glabrous; embryo straight or curved.
± 50 species: temperate western North America; southwestern South America. (Greek: divided flower, for perianth lobes) Involucre length measurements including teeth and awns unless otherwise specified.

Key to Chorizanthe

C. membranacea Benth. PINK SPINEFLOWER
NATIVE
Plant erect, 1–6(10) dm, hairy. Leaf: blades (1)1.5–5 cm, 0.1–0.3 cm wide, generally linear, tomentose abaxially. Inflorescence: bracts generally 2 (or a whorl of 3–5), awns 0.5–1 mm, straight; involucre 3–4 mm, 3-angled, 6-ribbed, tomentose to ± glabrous, ± swollen, teeth 5, 0.5–1 mm, margins scarious, awns 0.7–1.5 mm, hooked. Flower: 1(2), (1.5)2.5–3 mm, hairy; perianth white to rose, lobes entire; stamens 9. Fruit: 2.5–3 mm.
n=19–20,(22),40–41. Common. Sand, gravel or rocks; 40–1400 m. Northwestern California, Cascade Range, Sierra Nevada Foothills, Tehachapi Mountain Area, Great Central Valley, Central Western California, n Western Transverse Ranges; south-central Oregon. Apr–Jul [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 23 2014
Citation for this treatment: [Author of taxon treatment] 2013. Chorizanthe, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=19342, accessed on Oct 23 2014

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Geographic subdivisions indicated for the distribution of Chorizanthe membranacea Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.