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Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all ± alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1–3, less often many in circle; blade generally pinnate or ± palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
± 40 genera, 500 species: worldwide, especially dry areas. [Windham 1993 FNANM 2:122–186] Definition of Cheilanthes, related genera problematic; traditional limits often untenable. —Scientific Editors: Alan R. Smith, Thomas J. Rosatti.
Key to Pteridaceae
Plant in soil or rock crevices; rhizome short- to long-creeping[-decumbent], generally many-branched, scales generally lance-linear, pale to dark, mid-stripe dark or not. Leaf: < 75 cm, young leaf tip hooked or coiled; stipe cylindric, red-brown to ± black; blade generally 2–3-pinnate, generally oblong to narrowly triangular; segments generally small, ± flat or abaxially concave (from recurved margins). Sporangia: along margin, in discrete patches to continuous, partly to completely covered by recurved margin (generally not recurved in Cheilanthes cooperae).Key to Cheilanthes
150+ species: generally America, generally dry areas. (Greek: lip flower, from location of sporangia)
Rhizome short-creeping; scales pale with red-brown to ± black mid-stripe ± to margin or not. Leaf: 6–14(20) cm, 1.5–3 cm wide, dark green; stipe < 1 mm wide, scales lanceolate, ciliate at base, pale; blade 3-pinnate; segments small, ± round, abaxially concave, with pale to red-brown scales, scales < 1 mm wide, ciliate at base, covering more deeply dissected scales, barely exceeding segment margin, adaxially glabrous or with deeply dissected scales; young leaf tip hooked. Sporangia: generally obscured by scales, recurved segment margin.
Crevices, bases of rocks; 300–2800 m. Outer North Coast Ranges, High North Coast Ranges, High Sierra Nevada, San Francisco Bay Area, South Coast Ranges, Modoc Plateau, White and Inyo Mountains; Oregon, Nevada. Hybridizes with Cheilanthes covillei, Cheilanthes gracillima. Allotetraploid hybrid of Cheilanthes covillei and Cheilanthes gracillima. [Online Interchange]
Previous taxon: Cheilanthes gracillima
Next taxon: Cheilanthes newberryi
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Mar 9 2014
Citation for this treatment: [Author of taxon treatment] 2012. Cheilanthes, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=19078, accessed on Mar 9 2014
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