Jepson Herbarium
		University of California, Berkeley

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Indexes to all accepted names and synonyms: | A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |

Annual to shrub, tree [ vine]; monoecious or dioecious.
Stem: generally branched [ fleshy or spiny].
Leaf: generally simple, alternate or opposite, generally stipuled, petioled; blade entire, toothed, or palmately lobed.
Inflorescence: terminal or axillary panicle, raceme or spike, or (Chamaesyce, Euphorbia) a compact unit enclosed by an involucre appearing flower-like, terminal or axillary, 1 or in whorled, umbel-like, or cyme-like arrays.
Flower: unisexual, ± radial; sepals generally 3–5, free or fused; petals generally 0; stamens 1–many, free or filaments fused; ovary superior, chambers 1–4, styles free or fused, simple or lobed.
Fruit: generally capsule.
Seed: 1 per chamber; seed scar appendage sometimes present, pad- to dome-like.
218 genera, 6000+ species: ± worldwide especially tropics; some cultivated (Aleurites, tung oil; Euphorbia species; Hevea, rubber; Ricinus). Many species ± highly TOXIC. [Wurdack & Davis 2009 Amer J Bot 96:1551–1570] Eremocarpus moved to Croton. Tetracoccus moved to Picrodendraceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Webster 1994 Ann Missouri Bot Gard 81:1–144]

Key to Euphorbiaceae

Annual, perennial herb, glabrous to hairy; sap milky; generally monoecious.
Stem: prostrate to erect, < 5 dm; branches alternate.
Leaf: cauline, opposite, short-petioled; blade base generally asymmetric, veins dark green.
Inflorescence: flower-like, generally 1 per node; involucre ± bell-shaped, bracts 5, fused; glands 4, distal appendages generally colorful, petal-like; flowers central.
Staminate flower: 3–many, generally in 5 clusters around pistillate flower, each flower a stamen.
Pistillate flower: 1, central, stalked; ovary chambers 3, styles 3, separate or fused at base, divided to entire.
Fruit: capsule, round to 3-angled or -lobed in ×-section.
Seed: generally 4-angled, smooth or sculptured.
± 250 species: dry temperate, subtrop worldwide, especially Am. (Greek: ancient name for kind of prostrate plant) [Wheeler 1941 Rhodora 43:97–154, 168–286]

Perennial.
Stem: prostrate to erect, hairy.
Leaf: 2–10 mm; stipules generally separate, minute; blade ovate, acute, entire, hairy.
Inflorescence: involucre < 2 mm, urn-shaped, hairy; gland < 0.5 mm, ovate; appendage wider than gland, entire, white to pink.
Staminate flower: 5–10.
Pistillate flower: style divided 1/2.
Fruit: < 2 mm, spheric, lobed, hairy.
Seed: ± 1 mm, ovoid, transversely ridged, white to brown.
Sandy flats; < 300 m. Sonoran Desert;

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Next taxon: Chamaesyce fendleri

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.