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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

CAMISSONIA SUN CUP
Annual, from taproot; rosette generally ± 0. Leaf: cauline, alternate, simple, generally linear to narrowly elliptic. Inflorescence: bracted; spike or raceme, nodding in bud, erect in fruit, flowers only at distal nodes. Flower: radial, generally opening at dawn; sepals 4, reflexed singly or in pairs; petals 4, yellow, generally fading red, often with red basal spots; stamens 8, longer ones opposite sepals, anthers attached at middle, pollen grains 3-angled except in polyploid taxa, at 20×; ovary chambers 4, stigma hemispheric, generally > anthers and cross-pollinated, or ± = anthers and self-pollinated. Fruit: ± cylindric, straight to wavy, distorted by seeds at maturity, dehiscent throughout most of its length; pedicel ± 0 or <= 2(15) mm, 0 or shorter in flower. Seed: in 1 row per chamber, narrowly obovoid, smooth (minutely pitted), glossy.
12 species: western North America (especially California Floristic Province), 1 South America. (L.A. von Chamisso, French-born German botanist, 1781–1838) [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Polyploidy and self-pollination have predominated in evolution of genus. Not monophyletic as treated in TJM (1993); segregates moved to Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, Tetrapteron (Wagner et al. 2007).
Unabridged references: [Raven 1969 Contr US Natl Herb 37:161–396]

Key to Camissonia

C. benitensis P.H. Raven SAN BENITO EVENING-PRIMROSE
NATIVE
Slender; hairs spreading, of 2 types (linear, white; rod-shaped, transparent), in inflorescence also glandular. Stem: erect or decumbent, 3–20 cm, peeling; branches widely spreading, wiry. Leaf: 7–20 mm, narrowly elliptic, minutely serrate. Flower: hypanthium ± 1.2 mm; sepals 3.2–3.5 mm, remaining adherent in pairs; petals 3.5–4 mm, yellow fading ± red, basal spots 2; < 10% of pollen grains 4-angled. Fruit: 15–40 mm, 0.8–1.3 mm wide, slightly swollen by seeds, straight or wavy, ± sessile. Seed: 0.6–0.8 mm.
2n=28. Sandy or gravelly serpentine soil; ± 600–1200 m. Inner South Coast Ranges (Fresno, Monterey, San Benito cos.). Self-pollinated. Threatened by off-road vehicles. Apr–Jun [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 23 2014
Citation for this treatment: [Author of taxon treatment] 2013. Camissonia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=16897, accessed on Nov 23 2014

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click for enlargement Camissonia benitensis
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2006 Aaron Schusteff

Geographic subdivisions indicated for the distribution of Camissonia benitensis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.