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MONTIACEAE MINER'S LETTUCE FAMILY

John M. Miller, except as noted

Annual to perennial herb; generally fleshy. Stem: 1–many, generally glabrous. Leaf: simple, alternate or opposite. Inflorescence: axillary or terminal; cyme, raceme, panicle, umbel, or flower 1. Flower: bisexual, radial; sepals generally 2(9), free; petals (1)2–19, free or ± fused; stamens 1–many, epipetalous or not, anthers pink, rose, or yellow; ovary superior, chamber 1, ovules 1–many, placenta basal or free-central; styles (0)1–8, generally fused at base, branched. Fruit: capsule, circumscissile or 2–3-valved. Seed: 1–many, shiny or ± pebbly or sculptured, black or gray, generally with oil-filled appendage as food for ants.
± 22 genera, ± 230 species: generally temperate America, Asia, Australia, Europe, Kerguelen Is, New Zealand, southern Africa, poorly represented in Europe; some cultivated (Lewisia, Calandrinia). [Ogburn & Edwards 2009 Amer J Bot 96:391–408] Details of flowers, seeds require 20× magnification. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Applequist et al. 2006 Syst Bot 31:310–319; Nyffeler et al. 2008 Haseltonia 14: 26–36]
Unabridged note: May include Hectorellaceae.

Key to Montiaceae

CALYPTRIDIUM PUSSYPAWS

John M. Miller & C. Matt Guilliams

Annual, perennial herb, ± fleshy, from taproot or fibrous roots, glabrous. Stem: 1–several, generally spreading to ascending. Leaf: basal or basal and cauline, simple, oblanceolate to spoon-shaped; basal rosetted. Inflorescence: raceme, panicle, or umbel, scapose, bracts generally < sepals, leaf-like or not; flowers generally on 1 side of axis, persistent in fruit or not; pedicels ± jointed with a transverse groove or constriction. Flower: sepals 2, ovate to reniform, generally scarious or scarious-margined, persistent in fruit; petals 2–4, < sepals, tips adherent, forming cap in fruit (fruit cap), falling as 1 unit; stamens 1–3, anthers pink, rose, or yellow; style included to exserted, stigmas 2. Fruit: 2-valved, generally compressed, narrowly oblong to ± round, generally translucent, deciduous or not. Seed: 1–many, black, dull, fine- to coarse-papillate to shiny, smooth.
8 species: western North America. (Greek: cap, for petal tips in fruit) [Hershkovitz 2006 Gayana Bot 63:13–74]
Unabridged references: [Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz 1991a Ann Missouri Bot Gard 78:1009–1021; Hershkovitz 1991b Phytologia 70:209–225; Hershkovitz 1992 Syst Bot 17:220–238; Hershkovitz & Zimmer 2000 Molec Phylogen Evol 15:419–439; Hinton 1975 Brittonia 27:197–208; Howell 1945 Leafl W Bot 4:214–216; Kelley et al. 2003 FNANM: 4:460–464; Thomas 1956 Leafl W Bot 8:9–11]

Key to Calyptridium

C. monospermum Greene
NATIVE
Perennial herb < 50 cm; caudex short, thick; taproot slender to thick. Stem: generally spreading to ascending, leafy to not. Leaf: basal 1.5–6 cm, rosette 1; cauline generally present, 0.8–3 cm. Inflorescence: axillary, umbel, ± open (dense in small plants), >= 2 per rosette, 1–10 cm diam; pedicel ± 0 to short. Flower: sepals 3–8 mm, ± round, scarious; petals 4, 3–7 mm, rose to white; stamens 3, anthers pink to rose; style 2–4 mm, thread-like, exserted, falling with fruit cap. Fruit: 2–3.5 mm, widely ovate to ± round; deciduous or not. Seed: 1–4(8).
2n=44. Open areas, sandy or gravelly soils, conifer forest; 300–3970 m. Klamath Ranges, High North Coast Ranges, Cascade Range, Sierra Nevada, San Francisco Bay Area, Transverse Ranges, San Jacinto Mountains, Great Basin Floristic Province; southern Oregon, Nevada, northern Baja California. [Cistanthe monosperma (Greene) Hershk.] Hybridizes with Calyptridium umbellatum in Sierra Nevada. Apr–Sep [Online Interchange]

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Next taxon: Calyptridium parryi

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 17 2014
Citation for this treatment: [Author of taxon treatment] 2012. Calyptridium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=16846, accessed on Apr 17 2014

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Bioregions in which Calyptridium monospermum occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.