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CABOMBACEAE WATERSHIELD FAMILY

Thomas J. Rosatti

Perennial, aquatic, rhizomes in mud.
Stem: elongate, leafy, ends floating.
Leaf: of 2 kinds: submersed, palmately dissected, opposite [ whorled], short-petioled, not peltate; floating, simple, alternate, petioled, peltate, entire; stipules 0.
Inflorescence: flowers 1 on long axillary peduncles, generally on or above water surface.
Flower: bisexual, parts ± free; sepals, petals each 3[4], alternate, persistent; stamens 3–6 or 18–36(51), filaments slender, anthers opening lengthwise; carpels [1]2–18, ovaries superior, 1-chambered, ovules [1]2–3[5].
Fruit: achene- or follicle-like, indehiscent.
Seed: 1–3, aril 0.
2 genera, 6 species: temperate, tropical America, Africa, e Asia, Australia; some Cabomba species cultivated for aquaria. [Taylor & Osborn 2006 Amer J Bot 93:344–356] Separation from Nymphaeaceae (carpels fused, fruits berry-like) confirmed by molecular data (Les et al. 1999 Syst Bot 24:28–46). —Scientific Editors: Bruce G. Baldwin, Robert E. Preston.
Unabridged references: [Les et al. 1999 Syst Bot 24:28–46; Taylor & Osborn 2006 Amer J Bot 93:344–356; Wiersema 1997 FNANM 3:78–80]
Unabridged note: Pollinated by wind (Brasenia) or insects (Cabomba).

Key to Cabombaceae

BRASENIA WATERSHIELD

1 sp. (C. Brasen, Danish surgeon, plant collector, 1738–1774)
Unabridged etymology: (Christoph Brasen, 1738–1774, Danish surgeon and plant collector, leader of the 1771 missionary expedition that established the Moravian mission of Nain on the coast of Labrador (Charters & Hollombe, pers. comm.))

B. schreberi J.F. Gmel.
NATIVE
Submersed parts covered with thick mucilage.
Stem: slender, 3–20 dm, branching, red-brown.
Leaf: centrally peltate; blade << petiole, 3.5–12(13.5) cm, 2–8 cm wide, elliptic to ovate, ± entire.
Inflorescence: flowers 1 in axils.
Flower: perianth 10–20 mm, ± red-purple or purple-brown; petals ± > sepals, linear-oblong; filaments thread-like; carpels (2)4–18, stigmas linear- decurrent.
Fruit: 6–8(10) mm, oblong, leathery.
Seed: 1–2.
2n=80. Ponds, slow streams; < 2200 m. North Coast Ranges, High Cascade Range, High Sierra Nevada, Sacramento Valley, Modoc Plateau (except Warner Mountains); to Alaska, Montana; e North America, Central America, South America, Africa, e Asia, e Australia. Apr–Oct [Online Interchange] {CNPS list}
Unabridged synonyms: [Brasenia peltata Pursh; Brasenia purpurea (Michx.) Casp.; Cabomba peltata (Pursh) F. Muell.; Hydropeltis purpurea Michx.]

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.