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Perennial herb from caudex or rhizome, generally ± hairy. Stem: often ± leafy on proximal 1/2, rarely trailing and leafy throughout. Leaf: generally simple, basal and/or cauline, generally alternate, generally petioled; veins ± palmate. Inflorescence: raceme or panicle, generally ± scapose. Flower: generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens 3, , 5, 8, or 10; pistils 1 (carpels ± fused, ovary lobed, chambers 1 or 2, placentas generally 2(3), axile or parietal or occasionally proximally axile and distally marginal in ovary lobes) or 2 (carpels free, placentas marginal), ovary nearly superior to inferior, occasionally more superior in fruit, styles generally 2(3). Fruit: capsule (generally 2(3)-beaked, valves generally 2(3), generally equal) or 2 follicles. Seed: generally many, small.
± 30 genera, 600 species: especially northern temperate, arctic, alpine; some cultivated (Bergenia, Darmera, Heuchera, Saxifraga, Tellima, Tolmiea). [Soltis et al. 2001 Ann Missouri Bot Gard 88:669–693; Okuyama et al. 2008 Molec Phylogen Evol 46:560–575] California Mitella moved to Mitellastra, Ozomelis, Pectiantia; Suksdorfia ranunculifolia to Hemieva. Parnassia moved to Parnassiaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Soltis 1988 Syst Bot 13:64–72]
Key to Saxifragaceae
Plant glandular; rhizome scaly; bulblets 0. Leaf: basal and cauline, distal reduced, becoming bract-like; stipules inconspicuous to leaf-like; blade round to ovate, base cordate to reniform, occasionally truncate, obtuse, or tapered, primary lobes 3–many, shallow to deep, teeth sharp-tipped. Inflorescence: ± flat-topped or ± tapered to tip, > 15-flowered, generally 1-sided; bracts sessile to short-petioled, proximal leaf-like. Flower: hypanthium partly fused to ovary; petals often ephemeral; stamens 5; pistil 1, ovary > 1/2-inferior, chambers 2, placentas 2, axile. Fruit: capsule.Key to Boykinia
8 species: North America, Asia. (S. Boykin, Georgia naturalist, 1786–1848) [Gornall & Bohm 1985 Bot J Linn Soc 90:1–71]
Plant (2)5–1.3 dm, stout. Leaf: 9–50 cm; stipules conspicuous, ± 1 cm or more, green, entire or teeth generally sharp-tipped, distal leaf-like; petiole 5–35 cm; blade < 20 cm wide, divided > 1/2 to base, lobes and teeth ± straight-sided. Inflorescence: ± flat-topped; branches occasionally coiled at tip; flowers crowded. Flower: calyx lobes 2–5 mm, triangular to elliptic; petals 5–7 mm, elliptic to round.
Shaded, moist meadows, streambanks; < 2500 m. Klamath Ranges, n Outer North Coast Ranges (Grouse Mtn), n&c High Sierra Nevada; to Washington, Montana. Jun–Jul [Online Interchange]
Previous taxon: Boykinia
Next taxon: Boykinia occidentalis
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 23 2014
Citation for this treatment: [Author of taxon treatment] 2013. Boykinia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=16006, accessed on Jul 23 2014
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