Directory       News       Site Map       Home
         
    Jepson eFlora: Taxon page
Key to families | Table of families and genera

Previous taxon Indexes to all accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Previous taxon

OPHIOGLOSSACEAE ADDER'S-TONGUE FAMILY

Donald R. Farrar, except as noted

Perennial herb, small, fleshy, generally glabrous; caudex generally underground, unbranched; roots glabrous with bulblets or plantlets or not. Leaf: generally 1 per caudex per year, divided into 2 facing parts with a common stalk,(0)1 sterile and 1(2) fertile (fertile occasionally aborted); sterile photosynthetic part (trophophore) separated from spore-bearing part (sporophore) at to well above ground level; trophophore simple to compound, veins free and forked or netted with included veinlets; sporophore simple to compound, or 0 in young plants. Sporangia: dehiscent into 2 valves, ± 1 mm wide, thick-walled.
10 genera, 80–100 species: ± worldwide, generally rare or overlooked. [Hauk et al. 2003 Molec Phylogen Evol 28:131–151; Kato 1987 Gard Bull Straits Settlem 40:1–14] —Scientific Editors: Alan R. Smith, Bruce G. Baldwin, Thomas J. Rosatti.
Unabridged note: Distantly related to most (leptosporangiate) ferns. Haploid (gametophyte) generation underground. Both diploid and haploid generations obligately mycorrhizal. The family Psilotaceae (whisk ferns, 2 genera), sister to Ophioglossaceae, is represented in California (South Coast) by 1 (of 2 total) apparently introduced species, Psilotum nudum (L.) P. Beauv. Psilotum is easily distinguished by the dichotomously branching, almost leafless green stems, lack of roots, and large (2–3 mm) 3-lobed sporangia; sporangia are borne on the adaxial (upper) side of a minute (± 1 mm) forked leaf. Pantrop, subtrop (nearest native populations in Arizona and in Sonora, Mexico); expected in cultivation areas, especially at bases of old palms, possibly brought in on root masses as subterranean gametophytes. [Pryer et al. 2004 Amer J Bot 91:1582–1598]

Key to Ophioglossaceae

BOTRYCHIUM MOONWORT
Roots smooth, pale yellow, without bulblets or plantlets. Leaf: deciduous; bud glabrous; sporophore and trophophore (or 2 sporophores) joined at or well above ground level; trophophore generally 1–2-pinnate (simple or entire or 0), linear to deltate to ternately triangular, thin to fleshy, pinnae ovate to oblong and midribbed or wedge- to fan-shaped and not midribbed, veins free, forked; sporophore 1–2-pinnate, rarely absent. Sporangia: not sunken in axis; stalk 0 or short.
25–35 species: generally temperate to arctic or alpine. (Greek: bunch of grapes, from clusters of sporangia) [Stensvold 2007 Ph.D. Dissertation, Iowa State Univ; Wagner & Wagner 1993 FNANM 2:85–106] Difficult, needs study; most species uncommon, sporadic; good sampling of populations highly desirable in specimens, which must be carefully spread and pressed for identification. Botrychium multifidum moved to Sceptridium. Botrychium pedunculosum W.H. Wagner, differing from Botrychium pinnatum in having trophophore stalk ± = trophophore rachis (vs trophophore stalk 0 to 1/10 trophophore rachis), recently confirmed for California, based on discovery in summer of 2010 near Reynolds Creek, western of Yosemite National Park, Calaveras Co.

Key to Botrychium

B. lineare W.H. Wagner SLENDER MOONWORT
NATIVE
Leaf: sporophore, trophophore joined distal to mid-leaf; trophophore stalk 0–3(6) mm, blade 1-pinnate, 1–4 cm, < 1.5 cm wide, oblong to narrowly deltate-ovate, thin but firm, green to white-green, pinnae well separated, ascending, 3–6 pairs, linear or fan-shaped, deeply cleft into linear segments, not midribbed, side margins of simple pinnae or segments of cleft pinnae not converging or converging at < 20°, outer margins entire to coarsely and bluntly toothed, basal pinnae occasionally with scattered marginal sporangia; sporophore 1–2-pinnate, oblong to deltate, branches stiff, ascending, strongly overlapping, stalk 0.2–0.5 × trophophore, sporangia crowded.
2n=90. Moist meadows; 2500–4000 m. c&s High Sierra Nevada; to Alaska, eastern to Minnesota. [Online Interchange] {CNPS list}

Previous taxon: Botrychium crenulatum
Next taxon: Botrychium lunaria

Contact/Feedback

Name search

Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 19 2014
Citation for this treatment: [Author of taxon treatment] 2013. Botrychium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=15924, accessed on Sep 19 2014

Copyright © 2014 Regents of the University of California
We encourage links to these pages, but the content may not be downloaded for reposting, repackaging, redistributing, or sale in any form, without written permission from The Jepson Herbarium.


Geographic subdivisions indicated for the distribution of Botrychium lineare Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
READ ABOUT YELLOW FLAGS
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.