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SAXIFRAGACEAE SAXIFRAGE FAMILY

Michael S. Park & Patrick E. Elvander, except as noted

Perennial herb from caudex or rhizome, generally ± hairy. Stem: often ± leafy on proximal 1/2, rarely trailing and leafy throughout. Leaf: generally simple, basal and/or cauline, generally alternate, generally petioled; veins ± palmate. Inflorescence: raceme or panicle, generally ± scapose. Flower: generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens 3, [4], 5, 8, or 10; pistils 1 (carpels ± fused, ovary lobed, chambers 1 or 2, placentas generally 2(3), axile or parietal or occasionally proximally axile and distally marginal in ovary lobes) or 2 (carpels free, placentas marginal), ovary nearly superior to inferior, occasionally more superior in fruit, styles generally 2(3). Fruit: capsule (generally 2(3)-beaked, valves generally 2(3), generally equal) or 2 follicles. Seed: generally many, small.
± 30 genera, 600 species: especially northern temperate, arctic, alpine; some cultivated (Bergenia, Darmera, Heuchera, Saxifraga, Tellima, Tolmiea). [Soltis et al. 2001 Ann Missouri Bot Gard 88:669–693; Okuyama et al. 2008 Molec Phylogen Evol 46:560–575] California Mitella moved to Mitellastra, Ozomelis, Pectiantia; Suksdorfia ranunculifolia to Hemieva. Parnassia moved to Parnassiaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Soltis 1988 Syst Bot 13:64–72]

Key to Saxifragaceae

BOLANDRA
Caudex scaleless, bearing bulblets. Leaf: basal and cauline, basal and proximal petioled, cauline stipuled, distal sessile, reduced; blade ± round, base cordate to reniform, lobes 3–7, deep, teeth sharp. Inflorescence: panicle-like, glandular; flowers few; bracts sessile, proximal often clasping, leaf-like. Flower: hypanthium free from ovary; petals not clawed; stamens 5, opposite sepals; pistil 1, ovary superior, chambers 2, placentas 2, axile. Fruit: capsule.
2 species: Washington, Oregon, California. (H.N. Bolander, California botanist, 1831–1897) [Gornall & Bohm 1985 Bot J Linn Soc 90:1–71] Closely related to Suksdorfia and Hemieva.

B. californica A. Gray SIERRA BOLANDRA
NATIVE
Plant 15–60 cm. Leaf: petiole 3–10 cm; blade 1–5 cm wide. Inflorescence: pedicel 0.5–3 mm. Flower: calyx lobes 3–6 mm, ovate to triangular; petals 4–7 mm, narrowly triangular, long-tapered, 1/3–1/2 as wide as sepals, green, margin purple; stamens << calyx lobes, opposite sepals. Seed: brown, minutely papillate.
2n=14. Rock crevices, wet cliffs; 1000–3000 m. n&c High Sierra Nevada (El Dorado to Mariposa cos.). May–Aug [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 1 2014
Citation for this treatment: [Author of taxon treatment] 2013. Bolandra, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=15845, accessed on Sep 1 2014

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Geographic subdivisions indicated for the distribution of Bolandra californica Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.