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NYCTAGINACEAE FOUR O'CLOCK FAMILY

Andy Murdock, except as noted

Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally ± radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3–5, generally notched to ± lobed; stamens 1–many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to ± flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock). [Spellenberg 2003 FNANM 4:14–17] —Scientific Editor: Thomas J. Rosatti.

Key to Nyctaginaceae

BOERHAVIA SPIDERLING
Annual, perennial herb. Stem: prostrate to erect; internode often with sticky region. Leaf: petioled; blade 1–6 cm, paler beneath, often brown-dotted. Inflorescence: panicle-like, branches ending in umbel, spike-like raceme, or paired or solitary flowers; bracts 1–3, free, not forming involucre. Flower: closing by evening; perianth bell-shaped, tube < 5 mm, limb < 3 mm; stamens 1–5; stigma ± spheric, generally exserted. Fruit: < 4 mm, club-shaped; ribs 4–5; wings 0.
± 40 species: warm regions worldwide. (H. Boerhaave, Dutch botanist, 1668–1738) [Spellenberg 2003 FNANM 4:17–28]

Key to Boerhavia

B. diffusa L.
NATURALIZED
Perennial herb. Stem: decumbent to erect, < 1 m, ± glabrous. Leaf: blade broadly lanceolate to round; glabrous to minutely puberulent; Inflorescence: umbel of (2)3–5 flowers, bracts generally strongly reduced, not leaf-like. Flower: 1–1.5 mm; perianth red-purple (pale red). Fruit: 3–4.5 mm; stalked glands on ribs, less dense between; tip broadly conic.
Disturbed places, often in sand, gravel along rivers; < 50 m. Sacramento Valley (Yolo Co.); tropics, subtropics worldwide. ± all year [Online Interchange]

Previous taxon: Boerhavia coulteri var. palmeri
Next taxon: Boerhavia triquetra var. intermedia

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 19 2014
Citation for this treatment: [Author of taxon treatment] 2012. Boerhavia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=15817, accessed on Apr 19 2014

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Bioregions in which Boerhavia diffusa occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.