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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

ATHYSANUS
Annual; hairs simple, or short-stalked with 2–4-rays. Stem: simple or branched at base, leafy. Leaf: basal not rosetted, short petioled, entire or dentate; cauline short-petioled or sessile, base not lobed, entire or dentate. Inflorescence: 1-sided; bracts 0. Flower: sepals erect, early-deciduous, base not sac-like; petals < to > sepals or rudimentary, white. Fruit: silicle, reflexed, indehiscent or late-dehiscent, unsegmented, round or obovate to elliptic, flat parallel to septum, glabrous to hairy; pedicel recurved. Seed: 1 or 6–12, in 1 row, flat, not winged.
2 species: western North America, northwestern Mexico. (Greek: without fruit fringe)
Unabridged note: Jepson 1901 Flower W Calif:224 united Athysanus and Heterodraba under the former, thus establishing nomenclatural priority. The 2 genera are indistinguishable in every character except those in the key below. Molecular data support the placement of the combined genus near Draba and Arabis.

Key to Athysanus

A. unilateralis (M.E. Jones) Jeps.
NATIVE
Stem: hairy; branched near base, prostrate to ascending, (3)7–25(35) cm. Leaf: basal nearly sessile, 0.5–2.2 cm, oblanceolate to obovate or oblong, entire to each side 1-toothed; hairs 3–4-rayed; cauline 2–5, tapered at base. Flower: sepals 0.6–1 mm; petals 1.3–1.7 mm. Fruit: 3–5 mm, obovate to elliptic or round; style 0.1–0.2 mm; pedicel 1–2(3) mm. Seed: 0.9–1.2 mm, plump, oblong.
Uncommon. Grassy, open slopes, flats, clay soils, floodplains, gypsum-clay slopes; 100–900 m. Cascade Range Foothills, s Sierra Nevada Foothills, Tehachapi Mountain Area, Great Central Valley, South Coast Ranges; Oregon, Baja California. [Heterodraba unilateralis (M.E. Jones) Greene] Feb–May [Online Interchange]
Unabridged synonyms: [Draba unilateralis M.E. Jones]

Previous taxon: Athysanus pusillus
Next taxon: Aubrieta

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 19 2014
Citation for this treatment: [Author of taxon treatment] 2013. Athysanus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=15179, accessed on Sep 19 2014

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Geographic subdivisions indicated for the distribution of Athysanus unilateralis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.