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Margriet Wetherwax & Dieter H. Wilken, family description, key to genera

Annual, perennial herb, woody vine [shrub], occasionally aquatic. Leaf: generally basal and cauline, alternate or opposite, simple or compound; petioles at base generally flat, occasionally sheathing or stipule-like. Inflorescence: cyme, raceme, panicle, or flowers 1. Flower: generally bisexual, generally radial; sepals 3–6(20), free, early-deciduous or withering in fruit, generally green; petals 0–many, generally free; stamens generally 5–many, staminodes generally 0; pistils 1–many, ovary superior, chamber 1, style 0–1, generally ± persistent as beak, ovules 1–many. Fruit: achene, follicle, berry, ± utricle in Trautvetteria, in aggregate or not, 1–many-seeded.
± 60 genera, 1700 species: worldwide, especially northern temperate, tropical mountains; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Helleborus, Nigella). some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus). [Whittemore & Parfitt 1997 FNANM 3:85–271] Taxa of Isopyrum in TJM (1993) moved to Enemion; Kumlienia moved to Ranunculus. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Ranunculaceae


Scott Simono

Perennial herb from caudex, rhizome, or tuber. Leaf: basal, generally many, simple to compound, generally petioled; blade or leaflets lobed to dissected or not, margins entire or toothed; in flower or fruit withered or not. Inflorescence: terminal, flowers 1 or 2–7[9] in cymes; peduncle erect; pedicel elongated in fruit; involucre bracts sessile or stalked, generally in 1–2 whorls of 2–5[9], simple to compound, ± like leaves or leaflets in size, shape. Flower: receptacle elongated in fruit; sepals 5–10[27], petal-like; petals generally 0; stamens 10–200; pistils many, styles persistent as beaks. Fruit: achene.
± 150 species: arctic, temperate worldwide; some cultivated for ornamental. (Greek: flower shaken by wind) [Dutton et al. 1997 FNANM 3:139–155] Species with long, plumose styles sometimes placed in Pulsatilla.
Unabridged references: [Hoot et al. 1994 Syst Bot 19:169–200]

Key to Anemone

A. tuberosa Rydb.
Plant 10–30(40) cm; caudex slender, top with tuber. Leaf: 1–3(5); petiole 5–7 cm; blade 1–2 ternate, generally glabrous; leaflet margins ± lobed or dissected in distal 2/3; terminal leaflet (1.5)2–3(3.5) cm, 1–2(2.5) cm wide; ultimate segments 4–8(12) mm wide. Inflorescence: flowers 1–3(5); peduncle ± glabrous proximally, woolly distally; involucre bracts in (1)2 whorls of 3, ± sessile, simple, 1–2(3)-pinnately lobed or dissected, sparsely soft-hairy, margins irregularly minutely serrate; ultimate segments occasionally 1.5–2.5 mm wide. Flower: sepals 8–10, 10–14(20) mm, (2)3–5(6) mm wide, linear-oblong, pink to white, sparsely hairy; stamens 50–60. Fruit: body 2–3.5 mm, round, flat, densely woolly; pedicel (5)7–15(22) cm; beak ± 1.5 mm, straight, soft-puberulent; aggregate (15)20–30 mm, ellipsoid.
2n=16. Rocky slopes, ledges; 900–1900 m. e Desert Mountains, e Mojave Desert, ne Sonoran Desert; to Utah, Texas, northern Mexico. Apr–May [Online Interchange]

Previous taxon: Anemone oregana var. oregana
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 29 2015
Citation for this treatment: [Author of taxon treatment] 2013. Anemone, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 29 2015

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click for enlargement Anemone tuberosa
See CalPhotos for additional images
© 2003 James M. Andre

Geographic subdivisions indicated for the distribution of Anemone tuberosa Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.