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VERBENACEAE VERVAIN FAMILY

Dieter H. Wilken

Annual to shrub [tree], generally hairy. Leaf: cauline, opposite, generally toothed; stipules 0. Inflorescence: raceme, spike, or head, generally elongated in fruit; bract generally 1 per flower. Flower: bisexual; calyx generally 4–5-toothed; corolla 4–5-lobed, radial to bilateral, salverform to 2-lipped; stamens 4–5 (if 4, generally in unequal pairs), epipetalous; ovary superior, 2- or 4-lobed, generally 2- chambered, style 1, often with 2 unequal lobes, only 1 stigmatic, lateral. Fruit: 2 or 4 nutlets, drupe-like, or capsule.
± 31 genera, ± 920 species: especially America tropics. Some cultivated (Lantana, Verbena, Vitex); some weedy worldwide (Lantana); some used for wood (Tectona, teak). Avicennia included in Acanthaceae. [Marx et al. 2010 Amer J Bot 97:1647–1663] —Scientific Editor: Bruce G. Baldwin.

Key to Verbenaceae

ALOYSIA
Shrub, strong-smelling. Leaf: blade lanceolate to ovate. Inflorescence: raceme or spike, longer than wide. Flower: calyx 4-toothed; corolla 5-lobed, ± 2-lipped; stamens 4; ovary 2-chambered, ovules 2, style unlobed, stigma ± spheric, terminal. Fruit: nutlets 2.
± 35 species: America. Some used for food flavoring, tea. (Maria Louisa, princess of Asturias and, later, queen of Spain, 18th century)

A. wrightii (Torr.) Abrams WRIGHT'S BEEBRUSH
NATIVE
< 2 m, ± rounded. Stem: branches many; twigs brown, angles white. Leaf: petiole < 4 mm; blade 4–17 mm, ovate to ± round, crenate, abaxial face densely hairy. Inflorescence: spike 1.5–6 cm; axis densely and finely tomentose; bract slightly < calyx, lanceolate. Flower: calyx 1.5–3 mm, puberulent; corolla 2.5–3.5 mm, white, lobes rounded, upper 2 larger. Fruit: < 2 mm.
Rocky, often limestone, slopes, Joshua-tree or pinyon/juniper woodland; 900–1600 m. e&s Desert Mountains, e Sonoran Desert; to Texas, northern Mexico. Aug–Oct [Online Interchange] {CNPS list}
Unabridged synonyms: [Lippia wrightii Torr.]

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Next taxon: Lantana

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 20 2014
Citation for this treatment: [Author of taxon treatment] 2013. Aloysia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=12765, accessed on Oct 20 2014

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Geographic subdivisions indicated for the distribution of Aloysia wrightii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.