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Perennial herb, shrub, [tree], generally rhizomed, caudexed or not, glabrous, glaucous, or hairy. Stem: spreading to erect, branched or not. Leaf: simple, 1–3-ternate, or pinnately compound, basal and cauline, generally alternate, deciduous or evergreen, petioled, stipuled. Inflorescence: generally raceme, spike, or panicle, scapose, terminal, or axillary. Flower: generally bisexual, radial; sepals 6–18 or 0, generally in whorls of 3; petals generally 6, in 2 whorls of 3, or 0; stamens 6–12(13), free or fused at base, in 2 whorls or not, anthers dehiscent by flap-like valves or longitudinal slits; ovary superior, chamber 1, ovules generally 1–10, style 1 or 0, stigma flat or spheric. Fruit: berry, capsule, achene [follicle].
16 genera, ± 670 species: temperate, tropics worldwide; some cultivated (Berberis, Epimedium, Nandina (heavenly bamboo), Vancouveria). [Wang 2007 Syst Bot 32:731–742] Lower sepals sometimes called "bracteoles," inner petals "staminodes". —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Ernst 1964 J Arnold Arbor 45:1–35]
Key to Berberidaceae
Perennial herb; rhizome scaly; caudex short, erect; wood, especially of roots, often yellow. Leaf: 1–few, basal, long-petioled, 1-ternate; leaflets triangular to fan-shaped, bases tapered. Inflorescence: ± scapose, spike, dense; lateral flowers generally unisexual, terminal bisexual. Flower: perianth 0; stamens (8)9(13), anther valves flap-like, curled inward; ovule 1, style 0, stigma ± flat, furrowed. Fruit: achene, curved, furrowed, brown to red-purple.Key to Achlys
2 species: western North America, Japan. (Greek: thin mist or obscurity, from inconspicuous flowers)
Unabridged references: [Terabayashi 1981 Bot Mag (Tokyo) 94:141–157]
Plant 3–5 dm; rhizome internodes (7.5)9–10 cm. Inflorescence: < 4 cm. Fruit: 3.5–5 mm.
2n=24. Moist, shaded sites, conifer forest; < 1500 m. North Coast, w Klamath Ranges, w Outer North Coast Ranges; to British Columbia. Apr–Jun [Online Interchange]
Previous taxon: Achlys
Next taxon: Achlys triphylla subsp. triphylla
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 27 2015
Citation for this treatment: [Author of taxon treatment] 2013. Achlys, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=11785, accessed on Aug 27 2015
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|Geographic subdivisions indicated for the distribution of Achlys californica|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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