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NYCTAGINACEAE FOUR O'CLOCK FAMILY

Andy Murdock, except as noted

Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally ± radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3–5, generally notched to ± lobed; stamens 1–many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to ± flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock). [Spellenberg 2003 FNANM 4:14–17] —Scientific Editor: Thomas J. Rosatti.

Key to Nyctaginaceae

ABRONIA SAND-VERBENA
Annual, perennial herb, generally glandular. Stem: prostrate to ascending, generally ± red. Leaf: generally fleshy, petioled. Inflorescence: head or umbel; flowers maturing outer before inner or ± simultaneously; receptacle conic, ± smooth; bracts 5–10. Flower: perianth salverform to trumpet-shaped, generally fragrant, lobes 4–5; stamens 4–5, included; stigma linear, included. Fruit: body fusiform; wings (0)2–5, lobe-like, prominent, opaque, thick, not continuous above fruit body.
25 species: western North America. (Greek: graceful) [Galloway 2003 FNANM 4:61–69] Closely related to Tripterocalyx; relationships among species Abronia gracilis Benth. (Mexico), Abronia maritima, Abronia umbellata, and Abronia villosa need study; hybrids involve Abronia latifolia, Abronia maritima, Abronia umbellata; Abronia gracilis included in TJM (1993) based on misidentifications.

Key to Abronia

A. villosa S. Watson
NATIVE
Annual, glandular-hairy. Stem: prostrate to ascending, < 80 cm. Leaf: petiole 0.5–5 cm; blade 1–5 cm, 1–4.5 cm wide, triangular-ovate to ± round. Inflorescence: peduncle 2–10 cm; bracts 3–11 mm, lanceolate to narrowly ovate; flowers 15–35. Flower: perianth tube 1.3–3.5 cm, ± pink, throat with white spot, limb 6–18 mm wide, pale to bright magenta. Fruit: 5–10 mm, body with prominent raised veins, top with hardened beak (base of floral tube); wings 3–5, thin, rounded or angled, or 0. [Online Interchange]

Previous taxon: Abronia umbellata var. umbellata
Next taxon: Abronia villosa var. aurita

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Apr 23 2014
Citation for this treatment: [Author of taxon treatment] 2012. Abronia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=11568, accessed on Apr 23 2014

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click for enlargement Abronia villosa
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Charles Webber © 1998 California Academy of Sciences

Bioregions in which Abronia villosa occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.