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RUBIACEAE MADDER FAMILY

Robert E. Preston & Lauramay T. Dempster, except as noted

Annual to tree, vine.
Leaf: generally opposite ( whorled), entire; stipules generally fused to stem, adjacent pairs occasionally fused, or occasionally leaf-like and appearing like whorled leaves.
Inflorescence: cyme, panicle, spike, cluster, or flower 1, generally terminal and ± axillary.
Flower: generally bisexual; calyx ± 4(5)-lobed, occasionally 0 (Galium, Crucianella) or 6 (Sherardia); corolla generally radial, 4(5)-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1(2).
Fruit: drupe, berry, or 2 or 4 nutlets [ capsule].
± 500 genera, 6000 species: worldwide, especially tropics; many cultivated, including Coffea, coffee; Cinchona, quinine; many ornamental. [Robbrecht & Manen 2006 Syst & Geogr Plant 76:85–146] Diodia teres Walter doubtfully in CA. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Dempster 1979 Fl CA 4(2):1–47]

Key to Rubiaceae

CEPHALANTHUS BUTTON BUSH
Shrub, small tree.
Leaf: opposite or in whorls of 3; stipules not leaf-like.
Inflorescence: head, dense-spheric, peduncled.
Flower: calyx 4-lobed; corolla narrowly funnel-shaped; style thin, exserted, stigma ± spheric.
Fruit: nutlets 2–4, dry, brown, wider distally.
± 17 species: warm-temperate America, Asia, s Africa. (Greek: head flower)

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Next taxon: Cephalanthus occidentalis

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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