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CACTACEAE CACTUS FAMILY

Bruce D. Parfitt, except as noted

Perennial, shrub, tree, generally fleshy.
Stem: cylindric to spheric, or flat; surface smooth, tubercled, or ribbed (grooved); nodal areoles bearing flowers.
Leaf: generally 0 or early- deciduous, flat to ± cylindric.
Spines: areoles generally with central, radial spines, occasionally with glochids.
Flower: generally 1 per areole, bisexual [ unisexual], sessile, radial [ bilateral]; perianth parts generally many [5], scale-like to petal-like; stamens many; ovary inferior [superior], style 1, stigma lobes generally several [many].
Fruit: dry to fleshy or juicy, indehiscent to variously dehiscent, spiny, scaly, or naked; tubercled or smooth.
Seed: generally many, occasionally 0–few.
± 125 genera, ± 1800 species: Am (especially deserts), Africa; many cultivated, some edible. [Parfitt & Gibson 2004 FNANM 4:92–257] Spines smaller, fewer (0) in shade forms; yellow spines blacken in age. Introduced species increasingly escape cultivation. Hybridization common in some genera. Taxa of Escobaria in TJM (1993) moved to Coryphantha. —Scientific Editors: Bruce D. Parfitt, Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Hunt 2006 The New Cactus Lexicon, DH Books, Milborne Port, England]

Key to Cactaceae

MAMMILLARIA FISHHOOK CACTUS
Generally erect ( decumbent or prostrate), branched or not, branches 0–9(50).
Stem: 5–30 cm, [1.8]3–7.5[20] cm diam, spheric to cylindric [or obconic], firm to soft, not regularly segmented; ribs 0, tubercles prominent, conic to cylindric, not grooved.
Spines: [2]14–64(90) per areole, < 2 mm diam, needle-like [to hair-like or bristle-like], glabrous [or plumose], straight or hooked [or curved to crinkly]; central spines 1–4 [0–many] per areole, generally hooked.
Flower: lateral, in axils of tubercles, 1–5 [7.5] cm diam; perianth cream to white, pink, purple, or lavender; ovary glabrous, spines 0, scales 0.
Fruit: club-shaped or cylindric to ovoid [or barrel-shaped], indehiscent, generally red, spines 0.
Seed: 0.8–1.5 mm, generally shiny, generally pitted or raised-netted, black [brown to ± red or ± yellow], occasionally with aril.
150 species: North America. (Latin: nipple) [Hunt 1984 Bradleya 2:65–96; Hunt 1985 Bradleya 3:53–66; Hunt 1987 Bradleya 5:17–48]

Key to Mammillaria

Previous taxon: Grusonia pulchella
Next taxon: Mammillaria dioica

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Data provided by the participants of the Consortium of California Herbaria.
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