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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

SMELOWSKIA
[Annual] perennial herb; generally cespitose, caudex branched; hairs many-branched, tree-like, occasionally mixed with simple and stalked-forked hairs. Leaf: basal rosetted, 1- or 2-pinnately lobed or divided (entire); cauline petioled or sessile, base not lobed. Inflorescence: elongated or not; bracts 0 (present). Flower: sepal ascending to spreading, base not sac-like; petals spoon-shaped to obovate, white to pink or purple (yellow). Fruit: dehiscent, cylindric to 4-angled or flat parallel or perpendicular to septum, unsegmented; stigma entire or ± 2-lobed. Seed: 4–30, in 1 row, wingless.
27 species: western North America, e¢ral Asia. (T. Smielowsky, Russian botanist, 1769–1815) [Al-Shehbaz & Warwick 2006 Harvard Pap Bot 11:91–99]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 27 2014
Citation for this treatment: [Author of taxon treatment] 2013. Smelowskia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=10622, accessed on Jul 27 2014

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