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Key to families | Table of families and genera
This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press.
Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.
Annual, perennial herb, often ± glandular, especially above.
Stem: generally 1–4 mm wide in lower 1/2.
Leaf: palmately lobed or divided, ± hairy; segments generally palmately lobed and/or toothed; upper alternate or opposite; blade generally round in outline, base generally cordate.
Flower: radial (± bilateral); sepals ± awned to mucronate; stamens 10, outer 5 opposite petals (lacking anthers in G. pusillum), inner 5 alternate petals, nectary glands 5, alternate petals.
Fruit: mericarp generally ovoid, dehiscent, 1-seeded; basal callus generally present, often with long bristles.
± 400 species: temperate, tropical mountains. (Greek: crane, from fruit beak) [Aedo 2001 Anales Jard Bot Madrid 59:3–65; Aedo et al. 1998 Ann Missouri Bot Gard 85:594–630]
Unabridged references: [Aedo 2000–2001 Anales Jard Bot Madrid 58:39–82, 59:3–65; Aedo et al. 1998 Ann Missouri Bot Gard 85:594–630]
1. Perennial, from thick, scaly, underground caudex
2. Infl 1-flowered ..... G. potentilloides
2' Infl 2-flowered
3. Mericarp without basal callus ..... [G. palmatum; G. pyrenaicum]
3a. Petals 19–21 mm; leaf divided > 0.9 to base or midrib ..... G. palmatum [evidently known only from ± urban environments in CA]
3a' Petals 7–11 mm; leaf divided 0.5–0.6 to base ..... G. pyrenaicum [neither naturalized nor escaped in CA; collected only once, in 1914, on UC Berkeley campus]
3' Mericarp with basal callus
4. Petals glabrous adaxially, ciliate at base
5. Petals 12–21 mm; fruit beak 27–44 mm ..... G. oreganum
5' Petals 3–8.1 mm; fruit beak 8.8–15.1 mm
6. Pedicel hairs reflexed, appressed, 0.2–0.6 mm ..... G. core-core
6' Pedicel hairs spreading or reflexed, not appressed, 0.2–2 mm ..... G. solanderi
4' Petals 1/3–3/4 hairy adaxially, ciliate at base or not
7. Nectaries woolly abaxially; petals white; stigmas 3–4 mm ..... G. richardsonii
7' Nectaries glabrous abaxially, hair-tufted at top; petals white to red-purple; stigmas 3–8 mm
8. Fr narrow tip 5–6 mm, stigmas 6–8 mm; leaves 3–8 cm wide; petals 10–15 mm ..... G. californicum
8' Fr narrow tip 3–4 mm, stigmas 3–6 mm; leaves 7.1–16 cm wide; petals 17–20 mm ..... G. viscosissimum
1' Annual, biennial, from short, ± slender roots
9. Fertile stamens 5 (outer 5 lacking anthers) ..... G. pusillum
9' Fertile stamens 10 (inner 5 and outer 5 all with anthers)
10. Sepals with transverse flaps between lengthwise keels ..... G. lucidum
10' Sepals without flaps or keels
11. Mericarp ribbed or ± net-like or transversely wrinkled
12. Mericarp without collar-like rings or fibers at tip, transversely wrinkled ..... G. molle
12' Mericarp with 1–5 collar-like rings and long fibers at tip, ribbed or ± net-like
13. Petals 5–9.5 mm; mericarp with 3–5 collar-like rings at tip; anthers ± yellow ..... G. purpureum
13' Petals 10–14 mm; mericarp with 1–3 collar-like rings at tip; anthers ± purple ..... G. robertianum
11' Mericarp smooth (although sometimes ± ribbed or hairy or bristly)
14. Mericarp base with prong, without callus ..... G. dissectum
14' Mericarp base without prong, generally with callus
15. Mericarp glabrous except base generally ciliate ..... G. aequale
15' Mericarp ± hairy throughout except basal callus bristly
16. Fr beak with a narrow tip < 2 mm
17. Pedicel hairs spreading, glandular and nonglandular, sometimes also reflexed, not appressed, nonglandular; mericarp hairs dense, 0.5–1.8 mm ..... G. carolinianum
17' Pedicel hairs reflexed, appressed, nonglandular; mericarp hairs sparse, 0.2–0.7 mm ..... [G. texanum]
16' Fr beak with a narrow tip 2–6 mm
18. Peduncles without glandular hairs ..... G. columbinum
18' Peduncles with glandular hairs
19. Lf divided 0.85–0.95 to base ..... G. bicknellii
19' Lf divided 0.55–0.65 to base ..... G. rotundifolium
Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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