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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.



Alan R. Smith, except as noted

Perennial herb in soil or rock crevices; rhizome generally short-creeping, ascending, or erect, scales small to large, generally tan to brown, generally uniformly colored. Leaf: generally tufted or short-spaced, 5–200+ cm, generally ± alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade generally 1–3-pinnate, ± glabrous or with hairs, hair-like scales, or gland-tipped hairs on axes, veins generally free (or netted); rachis, costa generally grooved adaxially. Sporangia: sori round, oblong, J-shaped, or linear along veins; indusia 0 or oblong, J-shaped, reniform, or linear, or of many segmented hair- or scale-like fragments or lobes encircling sorus from below; spores elliptic, winged, ridged, or spiny, scar linear.
± 15 genera, 700 species: worldwide, especially tropics, wooded areas, but some genera (e.g., Cystopteris, Woodsia) generally temperate. See note, reference (Smith et al. 2006 Taxon 55:705–731) under Dryopteridaceae for removal of genera from that family to this. —Scientific Editor: Thomas J. Rosatti.

Key to Woodsiaceae

1. Leaves generally 30–150+ cm; stipe generally > 1.5 mm diam, if < then indusia 0; blade generally 25–130 cm, (5)10–60 cm wide; indusia 0, oblong, J-shaped, or reniform ..... ATHYRIUM

1' Leaves generally < 30 cm; stipe generally < 1.5 mm diam; blade generally 10–25 cm, 1–10 cm wide; indusia hood- or cup-like and of many segmented hair- or scale-like fragments or lobes

2. Indusium hood-like; pinnae sides unequal, acroscopic pinnules more spreading, larger, more incised; leaf blade axes abaxially glabrous ..... CYSTOPTERIS

2' Indusium cup-like, of many segmented hair- or scale-like fragments or lobes encircling sorus from below, often obscure in age; pinnae sides ± equal; leaf, blade axes abaxially with (without) short-stalked glands or gland-tipped hairs 0.1–1 mm ..... WOODSIA


Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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