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Key to families | Table of families and genera

This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press.

Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.

 

CHAMAESYCE PROSTRATE SPURGE

Daryl Koutnik


Annual, perennial herb, glabrous to hairy; sap milky; generally monoecious.
Stem: prostrate to erect, < 5 dm; branches alternate.
Leaf: cauline, opposite, short-petioled; blade base generally asymmetric, veins dark green.
Inflorescence: flower-like, generally 1 per node; involucre ± bell-shaped, bracts 5, fused; glands 4, distal appendages generally colorful, petal-like; flowers central.
Staminate flower: 3–many, generally in 5 clusters around pistillate flower, each flower a stamen.
Pistillate flower: 1, central, stalked; ovary chambers 3, styles 3, separate or fused at base, divided to entire.
Fruit: capsule, round to 3-angled or -lobed in ×-section.
Seed: generally 4-angled, smooth or sculptured.
± 250 species: dry temperate, subtrop worldwide, especially Am. (Greek: ancient name for kind of prostrate plant) [Wheeler 1941 Rhodora 43:97–154, 168–286]

Key to Chamaesyce

1. Infls generally in dense axillary cyme-like clusters, occasionally 1 per node; leaves 8–35 mm; stem generally erect (decumbent) ..... C. nutans

1' Infls 1 per node, occasionally crowded on lateral branches; leaves 1–28 mm; stem prostrate to ascending, sometimes erect

2. Involucre, fruit hairy; stem, leaves generally hairy or becoming glabrous

3. Perennial

4. Involucre urn-shaped; staminate flowers <= 10 — DSon ..... C. arizonica

4' Involucre ± bell-shaped; staminate flowers >= 15

5. Seed 3-angled, transversely 4–5-ridged, ridges rounded ..... C. pediculifera

5' Seed 4-angled, smooth to ± wrinkled

6. St and leaf hairs short, straight ..... C. polycarpa (2)

6' St and leaf tomentose or hairs long, appressed, dense

7. Involucre glands red ..... C. melanadenia

7' Involucre glands green to yellow ..... C. vallis-mortae

3' Annual

8. Staminate flowers 40–60; fruit 2–2.5 mm; n ScV ..... C. ocellata subsp. rattanii

8' Staminate flowers < 20; fruit <= 2 mm; widespread

9. Gland appendage deeply 3–5-lobed; involucre urn-shaped ..... C. setiloba

9' Gland appendage entire, shallowly scalloped, or 0; involucre bell-shaped or obconic

10. Gland appendage 0; seed smooth to slightly wrinkled ..... C. micromera (2)

10' Gland appendage present; seed wrinkled or ridged

11. Fr 1.5–2 mm; seed wrinkled; gland appendage narrower than gland ..... C. serpyllifolia subsp. hirtula

11' Fr <= 1.5 mm; seed transversely ridged or wrinkled; appendage equal to or wider than gland

12. Seeds transversely wrinkled; fruit evenly strigose; appendage equal in width to gland ..... C. maculata

12' Seeds transversely ridged; fruit hairy on lobes only; appendage equal or greater in width than gland ..... C. prostrata

2' Involucre, fruit glabrous; stem, leaves glabrous, sometimes hairy

13. Stipules fused into wide, membranous scale

14. Perennial; staminate flowers >= 15 ..... C. albomarginata

14' Annual; staminate flowers 5–10 ..... C. serpens

13' Stipules separate or fused, but not a wide, membranous scale

15. Perennial

16. Glands round, appendage 0; leaves 2–4 mm ..... C. parishii

16' Glands elliptic or oblong, appendage present; leaves 1–11 mm

17. Fr >= 2 mm; stipules separate; seed >= 2 mm ..... C. fendleri

17' Fr 1–1.5 mm; proximal stipules fused; seed 1–1.5 mm ..... C. polycarpa (2)

15' Annual

18. Lvs linear, base symmetric; stem prostrate to erect — PR, D

19. Glands 1–4, oval, appendage narrower than gland; fruit and seed > 1.5 mm; seed smooth; stem prostrate to ascending ..... C. parryi

19' Glands ± 4, round, appendage wider than gland or 0; fruit and seed <= 1.5 mm; seed transversely ridged; stem erect ..... C. revoluta

18' Lvs lanceolate to round, base asymmetric; stem prostrate to decumbent

20. Lf margin toothed, at least toward tip

21. Style entire; gland appendages deeply 3–5-lobed ..... C. hooveri

21' Style divided; gland appendages entire or shallowly lobed

22. Plant hairy to subglabrous; seed transversely ridged ..... C. abramsiana (2)

22' Plant glabrous; seed smooth, wrinkled, or transversely ridged

23. Seed transversely ridged; appendage wider than gland ..... C. glyptosperma

23' Seed smooth to wrinkled; appendage narrower than gland ..... C. serpyllifolia subsp. serpyllifolia

20' Lf margin entire

24. Seed ± 2 angled, flattened lengthwise, surface smooth; gland ovate ..... C. platysperma

24' Seed 3–4-angled, surface smooth, wrinkled, or transversely ridged; glands round or elliptic

25. Gland appendage present; seed transversely ridged ..... C. abramsiana (2)

25' Gland appendage 0; seed smooth or wrinkled

26. Fr < 1.5 mm; seed 1–1.5 mm; staminate flowers 2–5 ..... C. micromera (2)

26' Fr 2–2.5 mm; seed 1.5–2 mm; staminate flowers 40–60 ..... C. ocellata

27. Lf <= 15 mm, lanceolate to ovate; D ..... subsp. arenicola

27' Lf < 10 mm, ovate, sickle-shaped; CA-FP ..... subsp. ocellata

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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