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| Jepson eFlora
Key to families | Table of families and genera This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press. Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link. |
Perennial generally from short, thick, ± branched taproot; tuberous root generally 0 (or spheric).
Stem: prostrate to erect, scape-like or branched.
Leaf: generally in basal rosette and cauline, simple, entire or not; base wide; margin generally ± translucent.
Inflorescence: ± scapose; cyme, panicle, raceme, or ± umbel; stems 1–many, generally leafless but bracted, disjointing in age or not, 1–many flowered; pedicel 0–30 mm.
Flower: sepals 2–8, free, persistent; petals 4–19, white, cream, yellow, orange, pink, rose, purple, overlapped in bud, often with pink or dark purple veins; stamens 1–50; styles 2–8, fused at base, stigmas 2–8, thread-like.
Fruit: 6–9 mm, spheric or ovoid, circumscissile near middle or below, translucent.
Seed: 1–50, dark, generally shiny, smooth or finely tubercled, 1–4 mm in size.
18 species: w North America, 16 in flora. (Captain Meriwether Lewis, of Lewis & Clark Expedition, 1774–1809) [Wilson et al. 2005 W N Amer Naturalist 65:345–358] {hort link} Many hybrids, cultivated, including Lewisia ×whiteae Purdy in CA; Lewisia columbiana (A. Gray) B.L. Rob. not in CA.
Unabridged references: [Colley & Mineo 1985 Pacific Hort 46; Davidson 2000 Lewisias (Portland); Dempster 1996 Madroño 43:415–416; Elliott 1966 Bull Alpine Gard Soc Gr Brit 34:1–76; Foster, Carroll, & Hipkins 1997 Fremontia 25:15–19; Gankin & Hildreth 1968 Four Seasons 2(4):12–14; Heckard & Stebbins 1974 Brittonia 26:305–308; Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz & Hogan 2003 FNANM 4:476–485; Hohn 1975 Ph.D. Dissertation, Univ of Washington; Mathew 1989 Kew Magazine Monogr; Daubenmire 1975 Syesis 8:9–23]
Unabridged note: Many hybrids, cultivars (Mathew 1989), incl Lewisia ×whiteae Purdy in CA, as well as Lewisia columbiana (Howell ex A. Gray) B.L. Rob. from Douglas Co., OR north to BC; Lewisia maguirei A.H. Holmgren endemic to NV; Lewisia sacajaweana B.L. Wilson & E. Rey-Vizgirdas restricted to ID; Lewisia tweedyi, endemic to n Cascades of WA, BC now placed in Cistanthe.
1. Cauline leaves 3–5, whorled, less often 2, opposite; taproot 0; tuberous root spheric, with many rootlets ..... L. triphylla
1' Cauline leaves 0–many, opposite or alternate; taproot present; tuberous root 0
2. Basal leaves green after flower, not shriveled
3. Basal leaf blades linear to narrowly oblanceolate; leaf margin entire
4. Lvs flat or adaxialy grooved ..... L. ×whiteae
4' Lvs ± cylindric, adaxially not grooved ..... L. leeana
3' Basal leaf blades ovate, oblanceolate, ± spoon-shaped, or obovate to round (± linear); leaf margin entire or toothed
5. Petals 5–9 mm, white to pale pink with darker veins; inflorescence panicle, open
6. Pedicel 0.3–4 mm; petals 6–9 mm ..... L. cantelovii
6' Pedicel 3–8 mm; petals 5–6 mm ..... L. serrata
5' Petals (8)10–20 mm, generally pink-purple with pale and darker stripes, less often white, cream with pink-orange stripes, yellow, or orange; inflorescence cyme, ± umbel, or panicle, ± dense ..... L. cotyledon
7. Lf margin ± entire; petals (8)12–14 ..... var. cotyledon
7' Lf margin dentate or wavy; petals 12–20 mm
8. Lf margin dentate; 225–2200 m ..... var. heckneri
8' Lf margin ± wavy; 100–400 m ..... var. howellii
2' Basal leaves shriveled after flower, turning brown
9. Fls sessile; bract, sepal pairs resembling 4-parted calyx
10. Sepal margin entire, not glandular; petals 12–26 mm ..... L. brachycalyx
10' Sepal margin toothed, glandular or not; petals 10–30 mm ..... L. kelloggii
11. Lf blade > 4.5 cm, > 1 cm wide; petals >= 20 mm ..... subsp. hutchisonii
11' Lf blade < 4.5 cm, < 1 cm wide; petals <= 20 mm ..... subsp. kelloggii
9' Fls pedicelled; bract, sepal pairs not resembling 4-parted calyx
12. Fls breaking apart in fruit; sepals 2–9, scarious, margins entire to ± jagged, not toothed
13. Sepals 2; petals 5–9; proximal bracts 2–4 per node ..... L. disepala
13' Sepals (4)6–9; petals 10–19; proximal bracts 4–7(8) per node ..... L. rediviva
14. Sepals 10–12(15) mm; petals ± 15 mm; stamens 20–30 ..... var. minor
14' Sepals 15–25 mm; petals 18–35 mm; stamens 30–50 ..... var. rediviva
12' Fls not breaking apart in fruit; sepals 2, not scarious, margins entire, toothed, or gland-toothed
15. Cauline leaves opposite, not markedly smaller than basal ..... L. oppositifolia
15' Cauline leaves 0 or alternate, markedly smaller than basal
16. Basal leaves oblanceolate, spoon-shaped, or obovate; inflorescence 3–100-flowered
17. Cymes panicle-like, 20–100-flowered; petals 6–7, pale pink, base yellow-green; stamens 4–5 ..... L. congdonii
17' Cymes ± umbel-like, 3–11-flowered; petals 7–10, magenta or red, bases white or ± pink; stamens 10–13 ..... L. stebbinsii
16' Basal leaves thread-like to linear to narrow-lanceolate or -oblanceolate; inflorescence < 16-flowered
18. Fls 2.5–4 cm diam; petals >= 15 mm ..... L. longipetala
18' Fls 0.5–2 cm diam; petals <= 20 mm
19. Petals 9–20 mm; sepal margins entire to ± jagged, not gland-toothed (obscurely or irregularly toothed), tips ± acute ..... L. nevadensis
19' Petals 4–10 mm; sepal margins ± jagged, toothed, or gland-toothed (entire), tips ± rounded or truncate
20. Bract, sepal glands ± red or dark; stamens 6; stigmas 4 ..... L. glandulosa
20' Bract, sepal glands 0 or pale; stamens 4–8; stigmas 3–6 ..... L. pygmaea
Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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