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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.



Robert Patterson & Robyn Battaglia

Annual, perennial herb. Stem: generally erect, generally branched from base. Leaf: cauline, opposite, entire or lobes 3–9, palmate, linear to narrowly lanceolate or spoon-shaped, generally not fused by membrane. Inflorescence: head, open clusters, few-flowered cyme, or flower 1; bracts ± leaf-like, generally palmate-lobed, lobes generally not connected by translucent membrane; flowers sessile or not. Flower: sepals generally equal; corolla funnel-shaped, salverform, or bell-shaped, with hairy ring inside tube or generally not (determined at 10×); stamens attached at 1 level, pollen yellow.
30 species: western North America, Chile. (Greek: narrow tube, for corollas of some species) [Battaglia & Patterson 2001 Madroño 48:62–78] Calyx lobe membrane generally expressed as length relative to calyx or lobe length, or as width relative to calyx lobe.

Key to Leptosiphon

1. Perennial herb

2. Leaf lobes 0 ..... L. floribundus subsp. hallii

2' Leaf lobes palmate

3. Leaf lobes generally 3; pedicel generally 1–6 mm ..... L. floribundus

4. Leaf, calyx hairy ..... subsp. floribundus

4' Leaf, calyx glabrous ..... subsp. glaber

3' Leaf lobes generally 5; pedicel 0 or << 1 mm

5. Corolla tube > calyx; leaf lobes lance-linear; seeds > 2 mm ..... L. pachyphyllus

5' Corolla tube <= calyx; leaf lobes generally linear; seeds < 2 mm ..... L. nuttallii

6. Leaf, calyx sparsely hairy ..... subsp. nuttallii

6' Leaf, calyx densely hairy

7. Leaf lobes 3–7 mm; High North Coast Ranges ..... subsp. howellii

7' Leaf lobes 5–10 mm; East of Sierra Nevada ..... subsp. pubescens

1' Annual

8. Corolla not both salverform and in dense bracted heads

9. Flowers sessile

10. Corolla 15–30 mm; c Central Western California ..... L. grandiflorus

10' Corolla < 15 mm; s Southwestern California, Sonoran Desert ..... L. lemmonii

9' Flowers pedicelled

11. Hairy ring inside corolla 0, filaments glabrous; corolla generally < calyx

12. Stem branches generally above base or 0; corolla tube << lobes; stamens exserted; seeds 1 per chamber ..... L. harknessii

12' Stem branches at base; corolla tube >= lobes; stamens included; seeds > 1 per chamber ..... L. pygmaeus

13. Corolla white; Sierra Nevada Foothills, Great Central Valley, Central Western California, Southwestern California (except Channel Islands) ..... subsp. continentalis

13' Corolla blue; s Channel Islands (San Clemente) ..... subsp. pygmaeus

11' Hairy ring inside corolla present or, if 0, then filaments hairy; corolla > calyx

14. Hairy ring inside corolla 0; filaments hairy at base

15. Stem branched at base; calyx membrane as wide as lobes; corolla veins obscure ..... L. filipes

15' Stem branched above base; calyx membrane wider than lobes; corolla veins conspicuous ..... L. liniflorus

14' Hairy ring inside corolla present; filaments glabrous (rarely hairy at base in Leptosiphon septentrionalis)

16. Corolla 2–4 mm; stamens attached at or below hairy ring ..... L. septentrionalis

16' Corolla 4–15 mm; stamens attached above hairy ring

17. Corolla tube < lobes ..... L. aureus

18. Corolla lobes bright yellow ..... subsp. aureus

18' Corolla lobes white ..... subsp. decorus

17' Corolla tube > lobes

19. Stigmas 1–2 mm; corolla throat generally maroon; stamens attached at or above mid throat; San Joaquin Valley, San Francisco Bay Area, Inner South Coast Ranges ..... L. ambiguus

19' Stigmas <= 1 mm; corolla throat yellow; stamens attached at or below mid throat; Northwestern California, Cascade Range, n&c Sierra Nevada Foothills, n High Sierra Nevada, San Francisco Bay Area, Modoc Plateau

20. Corolla tube < 5 mm, white, pink, blue, or lilac, <= calyx ..... L. bolanderi

20' Corolla tube 10–15 mm, maroon, > 1.5 × calyx ..... L. rattanii

8' Corollas both salverform (tube >> calyx) and in dense, bracted heads

21. Calyx membrane at least as wide as lobes

22. Bracts white-ciliate

23. Corolla tube 10–25 mm, lobes 2–4 mm ..... L. ciliatus

23' Corolla tube 25–30 mm, lobes 5–8 mm ..... L. montanus

22' Bracts not white-ciliate

24. Leaf lobes oblanceolate; corolla lobes generally truncate or ± notched; s High Sierra Nevada ..... L. oblanceolatus

24' Leaf lobes linear to lance-linear; corolla lobes rounded; s High Sierra Nevada and elsewhere in s California

25. Bract lobes not connected by translucent membrane; corolla tube glabrous outside; San Gabriel Mountains, San Bernardino Mountains, Mojave Desert ..... L. breviculus

25' Bract lobes connected at base by translucent membrane; corolla tube hairy outside; s High Sierra Nevada, Tehachapi Mountain Area ..... L. nudatus

21' Calyx membrane much narrower than lobes

26. Corolla tube 7–8 mm, < 2 × calyx ..... L. serrulatus

26' Corolla tube generally > 10 mm, 2–6 × calyx

27. Calyx hairs sparse or 0 except on margins, nonglandular

28. Corolla lobes generally < 4 mm; stigmas <= 1(3) mm ..... L. bicolor

28' Corolla lobes > 4 mm; stigmas < or > 1 mm

29. Corolla lobes 4–6 mm; stigmas generally < 2 mm ..... L. jepsonii

29' Corolla lobes 6–14 mm; stigmas generally > 2 mm

30. Corolla lobes generally 6–9 mm, rounded; leaf lobes spoon-shaped, tips rounded ..... L. rosaceus

30' Corolla lobes generally > 8 mm, often abruptly soft-pointed; leaf lobes narrowly oblanceolate to linear, tips pointed ..... L. androsaceus

27' Calyx hairs dense, glandular or not

31. Leaf, calyx lobes needle-like; corolla lobes 3(5) mm, bright yellow ..... L. acicularis

31' Leaf, calyx lobes not needle-like; corolla lobes generally > 5 mm, yellow or otherwise colored, but not both < 5 mm and yellow

32. Corolla lobes 2–4 mm, tube 5–16 mm ..... L. minimus

32' Corolla lobes >= 4 mm, tube generally > 14 mm

33. Corolla lobes > 5 mm wide, rounded, bright yellow; calyx lobes generally deltate ..... L. croceus

33' Corolla lobes < 5 mm wide, not both rounded and yellow; calyx lobes generally narrowly lanceolate

34. Tip of middle leaf lobe rounded, generally > 2 mm wide at widest point; corolla lobes abruptly soft pointed or generally truncate, tube generally > 1 mm wide ..... L. latisectus

34' Tip of middle leaf lobe acute, generally < 2 mm wide at widest point; corolla lobes rounded, tube generally < 1 mm wide ..... L. parviflorus


Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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