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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.

 

HARMONIA

Bruce G. Baldwin


Annual 0.5–4 dm. Stem: erect. Leaf: proximal opposite, distal alternate, sessile, linear, entire or toothed, generally coarsely hairy, sometimes also minutely stalked-glandular, glands generally black, sometimes ± yellow. Inflorescence: heads radiate, 1 or in loose, ± umbel-like to flat-topped clusters; involucre obovoid to obconic, 2–5+ mm diam; phyllaries 3–8 in 1 series, lanceolate to oblanceolate, each enclosing a ray ovary, falling with fruit; receptacle flat to convex, glabrous or minutely bristly, paleae in 1 series between ray and disk flowers, free or fused, phyllary-like (more scarious), deciduous. Ray flower: generally 3–8; corolla bright yellow, ray fan-shaped to obovate. Disk flower: 7–30, bisexual or staminate, sometimes in same head; corolla bright yellow, tube < throat, lobes deltate; anthers ± yellow to ± brown, tips ovate-dentate to hemispheric; styles glabrous proximal to branches, tips awl-shaped, densely bristly. Fruit: ray fruit black, round in ×-section to ± compressed, weakly arched, bowed abaxially or not, beaked or beakless, glabrous, pappus 0 or of 3–12, lanceolate to awl-shaped, fringed to plumose scales; disk fruit black, ± club-shaped, ± round, glabrous or hairy, pappus of 7–11, generally linear to lanceolate or awl-shaped, fringed or plumose scales.
5 species: northwestern California. (Harvey Monroe Hall, California botanist, 1874–1932) [Baldwin & Strother 2006 FNANM 21:297–299]
Unabridged disk flower: anther bases sagittate.
Unabridged references: [Baldwin 2001 Madroño 48:293–297]

Key to Harmonia

1. Heads generally nodding in bud and fruit; ray pappus 0; disk pappus of lanceolate, long-tapered, minutely fringed scales 2–3.7 mm ..... H. nutans

1' Heads generally erect in bud and fruit; ray pappus 0.2–1.5 mm; disk pappus of linear to lanceolate or awl-shaped, or oblong to square, fringed or plumose scales 0.2–3.5 mm

2. Leaves ± evenly distributed on stem; ray fruit strongly bowed abaxially, beaked, beak 0.4–0.5 mm; disk flowers staminate ..... H. doris-nilesiae

2' Leaves unevenly distributed, mostly along main, central stem and immediately proximal to branches supporting heads; ray fruit slightly or not bowed abaxially, beak 0 or pappus elevated adaxially 0.1–0.2 mm; some or all disk flowers bisexual

3. Phyllaries densely white-soft-hairy near folded edges; ray fruit slightly bowed abaxially; disk pappus of linear and long-tapered to awl-shaped, plumose scales 1.2–3.5 mm ..... H. stebbinsii

3' Phyllaries coarsely hairy, or minutely so, near folded edges; ray fruit not bowed abaxially; disk pappus of lanceolate to linear or oblong to square, fringed scales 0.2–0.8 mm

4. Proximal, unbranched part of main, central stem generally > branches supporting heads, distal leaves of main stem not densely clustered; disk pappus of lanceolate to linear scales 0.6–0.8 mm ..... H. guggolziorum

4' Proximal, unbranched part of main, central stem generally < branches supporting heads, distal leaves of main stem densely clustered; disk pappus of oblong or square scales 0.2–0.5 mm ..... H. hallii

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Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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