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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.


Plant in soil or rock crevices; rhizome short-creeping-decumbent, generally 3–5(8) mm wide, scales lance-linear, mid-stripe dark. Leaf: stipe 5–20(32) cm, 0.5–2(3) mm wide; blade generally 2–3-pinnate, 2–8(15) cm, triangular or generally 5-sided, with white or yellow exudate abaxially, with exudate or not adaxially, main axis shallowly to deeply grooved adaxially; lowermost pinnae more strongly developed on basal side; veins free. Sporangia: along veins ± throughout; segment margins unmodified, recurved or not.
2 species: western North America. (Greek: 5 lines, for leaf blades) A puzzling complex of intergrading chemical, chromosomal, and morphological variants (see Yatskievych et al. 1990 Amer Fern J 80:9–17). {hort link}

Key to Pentagramma

1. Stipe dark brown to generally ± black; leaf blade adaxially with white exudate; Cascade Range Foothills, Sierra Nevada, Sacramento Valley(?), Inner South Coast Ranges ..... P. pallida

1' Stipe brown to red-brown; leaf blade adaxially generally without exudate; California Floristic Province, Great Basin Floristic Province, Desert Mountains ..... P. triangularis

2. Leaf blade adaxially not sticky, generally glabrous ..... subsp. triangularis

2' Leaf blade adaxially sticky or ± not, ± densely or sparsely glandular

3. Leaf blade adaxially ± not sticky, generally sparsely glandular; pinnules on basal side of lowermost pinnae deeply pinnately lobed to ± 1-pinnate ..... subsp. maxonii

3' Leaf blade adaxially generally sticky, ± densely glandular; pinnules on basal side of lowermost pinnae ± entire to shallowly pinnately lobed ..... subsp. viscosa


Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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