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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.



Barbara Ertter

Perennial herb, hairs short, simple, nonglandular and/or long, cross-walled, glandular. Stem: ± erect, from ± branched caudex or rhizomes. Leaf: basal and cauline, alternate, odd-1-pinnately compound; leaflets toothed, terminal generally >= lateral. Inflorescence: cyme, ± open; pedicels straight, bractlets 0. Flower: hypanthium ± shallow, bractlets 5; sepals ± triangular; petals < to > sepals, white to yellow; stamens generally 20–25, pollen sac 1, horseshoe-shaped; pistils many, styles fusiform, attached below fruit middle. Fruit: achene, glabrous.
30 species: northern temperate. (Greek: wood beauty) [Ertter 2007 J Bot Res Inst Texas 1:31–46] Recognition based on morphological, molecular evidence. Drymocallis ashlandica (Green) Rydb. (inflorescence narrow, petioles glandular, petals yellow) in southwestern Oregon, possibly northwestern California.
Unabridged note: Convergence of morphological, molecular evidence mandates recognition of Drymocallis; infrageneric taxonomy provisional. Key characters include relative proportions of 2 hair types: short simple nonglandular hairs and longer cross-walled glandular hairs.

Key to Drymocallis

1. Flowers opening narrowly, petals ± erect; styles 1.5–3 mm, ± slender

2. Petiole generally densely hairy with nonglandular hairs; terminal leaflet generally > 10-toothed, tip obtuse to rounded; n California ..... D. rhomboidea

2' Petiole sparsely hairy with glandular and nonglandular hairs; terminal leaflet < 10-toothed, tip ± truncate; Transverse Ranges ..... D. cuneifolia

3. Stem 20–45 cm; leaves 5–15 cm; San Bernardino Mountains ..... var. cuneifolia

3' Stem generally 5–20 cm; leaves 2–10 cm; San Gabriel Mountains ..... var. ewanii

1' Flowers opening widely, petals spreading; styles generally < 1.5 mm, swollen

4. Petals ± <= sepals, ± elliptic-(ob)ovate; stem and pedicel hairs generally glandular ..... D. glandulosa

5. Petals ± elliptic-ovate, 3.5–6 mm wide, cream to pale yellow; inflorescence appearing ± leafy (bracts ± 1/2 subtended branches); pedicels 1–5 mm

6. Petals generally 3.5–5 mm, generally pale yellow, sometimes cream; inland ..... var. glandulosa

6' Petals generally 4.5–6.5 mm, cream (± yellow); coastal ..... var. wrangelliana

5' Petals narrow-elliptic-obovate, 2–3.5 mm wide, yellow; inflorescence not appearing leafy (bracts < 1/2 subtended branches); pedicels 2–10 mm

7. Inflorescence branches spreading (angle generally 25–55°); leaflet teeth mostly double; sepal tip generally obtuse ..... var. reflexa

7' Inflorescence branches ascending (angle generally 10–30°); leaflet teeth mostly single; sepal tip acute ..... var. viscida

4' Petals > sepals, ± obovate to round; stem and pedicel hairs glandular or not — > 900 m

8. Stem generally 5–25 cm; sheathing leaf base generally appressed-hairy — generally glandular above; rocky areas, 2300–3900 m ..... D. pseudorupestris

9. Leaflets generally 4 per side; calyx with prominent nonglandular hairs 1–1.5 mm; styles generally ± red ..... var. crumiana

9' Leaflets generally 3 per side; calyx with nonglandular hairs generally < 1 mm or 0; styles generally golden-brown ..... var. saxicola

8' Stem generally 10–90 cm; sheathing leaf base generally glabrous

10. Inflorescence open, branches spreading (angle generally 20–40°); petals ± yellow; pedicel hairs glandular; n California ..... D. lactea var. austiniae

10' Inflorescence congested or narrow, branches ascending (angle generally 10–30°); petals cream or ± yellow; pedicel hairs nonglandular; Cascade Range, Sierra Nevada, Transverse Ranges, San Jacinto Mountains, East of Sierra Nevada

11. St-base hairs generally 2–3 mm, glandular; stem 40–90 cm; leaflet teeth generally double; 1200–2200 m ..... D. hansenii

11' St-base hairs < 1 mm, nonglandular; stem generally 20–60 cm; leaflet teeth generally single; 1800–3700 m ..... D. lactea var. lactea


Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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