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Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.

 

CALYPTRIDIUM PUSSYPAWS

C. Matt Guilliams & John M. Miller


Annual, perennial herb, ± fleshy, from taproot or fibrous roots, glabrous. Stem: 1–several, generally spreading to ascending. Leaf: basal or basal and cauline, simple, oblanceolate to spoon-shaped; basal rosetted. Inflorescence: raceme, panicle, or umbel, scapose, bracts generally < sepals, leaf-like or not; flowers generally on 1 side of axis, persistent in fruit or not; pedicels appearing jointed at base with a transverse groove or constriction or not. Flower: sepals 2, ovate to reniform, generally scarious or scarious-margined, persistent in fruit; petals 2–4, < sepals, tips adherent, forming cap in fruit (fruit cap), falling as 1 unit; stamens 1–3, anthers pink, rose, or yellow; style included to exserted, stigmas 2. Fruit: 2-valved, generally compressed, narrowly oblong to ± round, generally translucent, deciduous or not. Seed: 1–many, black, dull, with fine to coarse papillae, to shiny, with papillae 0.
9 species: western North America. (Greek: cap, for petal tips in fruit) [Hershkovitz 2006 Gayana Bot 63:13–74; Simpson et al. 2010 Madroño: 57:145–160] Calyptridium parryi var. arizonicum raised to species rank, as Calyptridium arizonicum, by Simpson et al. (2010) based on morphological and geographic distinctions.
Unabridged references: [Hershkovitz 1990 Phytologia 68:267–270; Hershkovitz 1991a Ann Missouri Bot Gard 78:1009–1021; Hershkovitz 1991b Phytologia 70:209–225; Hershkovitz 1992 Syst Bot 17:220–238; Hershkovitz & Zimmer 2000 Molec Phylogen Evol 15:419–439; Hinton 1975 Brittonia 27:197–208; Howell 1945 Leafl W Bot 4:214–216; Kelley et al. 2003 FNANM: 4:460–464; Thomas 1956 Leafl W Bot 8:9–11]

Key to Calyptridium

1. Perennial herb; style thread-like, 2–4 mm, exserted

2. Inflorescence >= 2 per rosette; cauline leaves generally present; petals rose to white ..... C. monospermum

2' Inflorescence generally 1 per rosette; cauline leaves generally 0; petals white ..... C. umbellatum

1' Annual; style thread-like or not, generally < 2 mm, generally included

3. Style 1–2 mm; seeds 1–3, 0.7–0.9 mm; c Sierra Nevada ..... C. pulchellum

3' Style < 0.5 mm; seeds >= 4, 0.4–0.8 mm; widespread, including e slope High Sierra Nevada (for C. roseum) but otherwise excluding c Sierra Nevada (except Calyptridium pygmaeum is in c&s High Sierra Nevada)

4. Seed < 0.5 mm, papillae generally 0 or marginal; pedicel 1–3 mm, slender

5. Petals 4; stamens generally 3 ..... C. pygmaeum

5' Petals 2; stamen 1 ..... C. roseum

4' Seed >= 0.5 mm, papillae marginal or throughout (except 0 in Calyptridium arizonicum); pedicel < 2 mm, thickened (slender, 1–2 mm in Calyptridium quadripetalum)

6. Petals generally 3; stamen 1; fruit oblong to linear; seeds 4–10, generally from base to above middle of fruit, often in ± 1 row ..... C. monandrum

6' Petals 4; stamens 1–3; fruit widely ovate or lanceolate to elliptic or oblong; seeds 5–many, generally from base to middle of fruit, not in ± 1 row

7. Seed papillae coarse, generally throughout; basal leaves generally persistent in fruit; flowers deciduous in fruit; sepals scarious throughout ..... C. quadripetalum

7' Seed papillae 0 or ± fine, throughout or marginal; basal leaves generally withering in fruit; flowers persistent in fruit to not; sepals generally scarious on margin or becoming scarious throughout

8. Seed 0.7–0.8 mm, papillae 0; fruit 6–7 mm; w Sonoran Desert ..... C. arizonicum

8' Seed generally 0.5–0.7 mm, papillae throughout or marginal; fruit < 6 mm; sw San Francisco Bay Area, s Outer South Coast Ranges, Transverse Ranges, San Jacinto Mountains, East of Sierra Nevada, Desert Mountains ..... C. parryi

9. Seed papillae throughout; s Outer South Coast Ranges, Transverse Ranges, San Jacinto Mountains ..... var. parryi

9' Seed papillae marginal; sw San Francisco Bay Area, East of Sierra Nevada, Desert Mountains

10. Seed 0.5–0.6 mm; sepals ovate, with generally narrowly scarious margin; 600–1050 m, sw San Francisco Bay Area ..... var. hesseae

10' Seed 0.6–0.7 mm; sepals ovate to reniform, with widely scarious margin or scarious throughout; 1500–2990 m, East of Sierra Nevada, Desert Mountains ..... var. nevadense

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Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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